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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />, <br /> <br />2 <br /> <br />of individual plasticity (Sousa 1984; Poffand Ward 1990; Schlosser 1991). <br /> <br /> <br />Due to high commercial and recreational value, some of the best documented and <br /> <br /> <br />studied examples of species' decline resulting from human-induced impacts are in fishes <br /> <br /> <br />such as salmons (Netboy 1974; Clay 1995). Overfishing, habitat loss or modification, <br /> <br /> <br />and introduced species all are implicated. Often more than one factor may come into play <br /> <br /> <br />at once, making it difficult to determine which is more important (Karr and Dudley <br /> <br /> <br />1989). More than a billion dollars have been spent to restore Columbia River salmon <br /> <br /> <br />fisheries yet the populations are rapidly approaching non-viable levels (Nehlsen et al. <br /> <br /> <br />1991). The problem, however, extends far beyond commerce and recreation. There are <br /> <br /> <br />few if any major rivers worldwide where one to many indigenous species are not <br /> <br /> <br />imperiled. All larger north temperate and tropical streams and rivers have been modified <br /> <br /> <br />(Sparks et al. 1990; Dynesius and Nilsson 1994; We1comme 1995). As of 1976 there <br /> <br /> <br />were 1426 hydropower and 27 pump-storage facilities and many smaller facilities <br /> <br />damming rivers and streams in the United States alone (USFPRC 1976; USFWS 1976). <br /> <br /> <br />The high human demand for water, particularly in arid lands, amplifies declines in aquatic <br /> <br /> <br />ecosystems (Walker et al. 1995). Vast changes in biotic structure, including species' and <br /> <br /> <br />community extinctions, have occurred and are under way (Miller 1961; Minckley and <br /> <br /> <br />Deacon 1968, 1991; Minckley 1991; Minckley and Douglas 1991). <br /> <br /> <br />For fishes in general, declining populations in altered aquatic habitats most often <br /> <br /> <br />result from shifts in the pattern of survivorship among younger cohorts (Houde 1987). <br /> <br /> <br />Declining abundance, particularly in fishes, provides a good subject for study because <br /> <br /> <br />extrinsic, density independent factors typically affect early survivorship (Lawler 1965; <br /> <br />-- <br />