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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br /> <br />CHAPTER 1. OVERVIEW AND STATEMENT OF THE PROBLEM <br /> <br /> <br />I. Introduction <br /> <br /> <br />1. General Review <br /> <br /> <br />Alteration and destruction of natural ecosystems are resulting from ever- <br /> <br />increasing human demands on the global environment (Brossard 1991; Lubchenco et al. <br /> <br /> <br />1991). The planet is no longer comprised of a series of pristine ecological systems, but <br /> <br /> <br />of systems artificialized in one way or another by human technology, some more <br /> <br /> <br />significantly than others (Pimm 1986). Human actions superimpose pervasive, continual <br /> <br />perturbation on natural disturbance regimes that maintain habitats and biotic communities <br /> <br /> <br />(Stanford et al. 1995). The result is suppression and loss of environmental heterogeneity <br /> <br /> <br />and a reduction in the productive abilities of biotic resources. Extinction rates have <br /> <br /> <br />increased, almost exponentially as a result of such actions (Myers 1986). Declines and <br /> <br /> <br />disappearances of species are correlated with physical alterations such as loss of breeding <br /> <br /> <br />sites, changes in habitat quality such as shifts of food availability or timing of abiotic <br /> <br /> <br />events, and changes in biotic pressures due to range extensions and introductions of <br /> <br /> <br />species better adapted to new conditions. <br /> <br />Life histories of species represent unique evolutionary solutions to age- or size- <br /> <br /> <br />specific patterns of survivorship (Tanner 1966, 1971; Wilbur et al. 1974; Southwood <br /> <br /> <br />1977; Reznick and Endler 1982). Species with broad tolerances, adjustable reproductive <br /> <br /> <br />strategies, and opportunistic tendencies may thus persist and even increase in the face of <br /> <br /> <br />change (Angermier 1985). Those with more constrained life history requirements will <br /> <br /> <br />likely decline, particularly when changes impinge on critical life-history stages (Boesch et <br /> <br /> <br />al. 1976; Poffand Ward 1990). Extinction may result when disturbance exceeds the limits <br /> <br />~f <br />