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<br />7 <br />LITERATURE REV1EW <br /> <br />General trends in abundance of nonnative fishes <br /> <br />Distributions of fishes introduced into the upper basin were shown on maps provided <br />by Tyus et al. (1982) and further discussed by Carlson and Muth (1989). Documented <br />reports of decreasing numbers of native and increasing numbers of nonnative fish are <br />numerous; however, it is unclear whether direct cause and effect relationships exist. Several <br />examples follow that identify an increase of nonnative fishes concurrently with a decrease <br />of native fishes <br /> <br />In three study areas on the upper Colorado and lower Gunnison rivers, Colorado, <br />Osmundson and Kaeding (1989a) noted that young native fishes progressively decreased in <br />abundance from 1986 through 1988, while introduced fishes, especially red shiner, fathead <br />minnow, and larval sand shiner, progressively increased in abundance. These trends in <br />abundance may have been related to annual differences in flow, i.e., 1986 was identified as <br />a high flow year, 1987 an average flow year, and 1988 a low flow year. <br /> <br />Behnke (1980) noted that an increase in redside shiner abundance occurred <br />concurrently with the decline of Colorado squawfish in the Green and Yampa rivers, Utah <br />and Colorado, and suggested that redside shiner had caused the decline. Holden (1977) <br />disagreed with this conclusion by noting that redside shiner were abundant in the 1960s when <br />Colorado squawfish reproduction was apparently successful. <br /> <br />The introduction and increase of northern pike and walleye in the Green River, Utah <br />was documented by Tyus and Beard (1990). Originally introduced into a Yampa River <br />tributary reservoir in 1977, northern pike invaded the Green River by 1981. Since that time, <br />they have increased in both range and abundance. Walleye were presumably established in <br />the Green River drainage in the 19605 by fish stockings in Duchesne River, Utah reservoirs. <br />Although widely distributed in the Green River, their numbers remain low but stable (Tyus <br />and Beard 1990). <br /> <br />The Yampa River provides another example of the abundance and distribution of <br />nonnative fish species. Miller et al. (1982b) studied the lower 206 kilometers (km) of the <br />Yampa River to its confluence with the Green River. Of 24 fishes collected, 15 were <br />introduced. Native fishes were more common within Yampa Canyon in Dinosaur National <br />Monument than other areas. Species diversity increased upstream as more nonnative fishes <br />were collected. They disagreed with Joseph et al. (1977) who implied that lack of <br />documented Colorado squawfish reproduction since 1969 was correlated with increasing <br />numbers of redside shiner, the dominant species in 1975-1976. Implications made by Joseph <br />et a!. (1977) were questioned because all collections were made in upper reaches, upstream <br />of documented Colorado squawfish reproduction (Miller et al. 1982b). The most abundant <br />species collected in Dinosaur National Monument by electrofishing and angling between <br />1987-1989, were common carp and channel catfish (Karp and Tyus 1mb). <br /> <br />I <br />