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<br />18 <br /> <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />DISCUSSION <br /> <br />Discharge, water temperature, and water quality <br /> <br />The 1 995 water year exemplified the seasonal nature and extreme variability <br />of the Little Snake River. Annual runoff volume in 1995 was 1.5 times higher than <br />the average of 415,200 acre feet lCrowfoot et al. 1996) and three times wetter <br />than the previous year which discharged about half the average volume (Ugland et <br />aJ. 1995). The unregulated characteristics of the river and the nature of the <br />hydrograph in 1995 were represented by the extreme peak to base-flow ratio <br />(165: 11 primarily caused by extremely low base flows which are typical for the <br />Little Snake River. Low baseflow dewatered all reaches but was most evident in <br />the sandy and braided middle reach. Fish habitat was maintained in the lower and <br />upper reaches by scattered refugia pools, connected with minimal surface flows <br />that were very shallow and created a barrier to fish movement between pools. <br />Shallow, widely dispersed flows also had a tremendous influence on water <br />temperature and caused diel variations of a greater magnitude than those in the <br />Yampa River. These and other physical influences likely caused a significant, but <br />unknown, role in initially attracting and eventually repelling humpback chub and <br />Colorado pikeminnow and structuring the fish community of the Little Snake River. <br /> <br />Fish community sampling <br /> <br />Humpback chub and Colorado pikeminnow occupied the lower 15 km of the <br />Little Snake River during summer runoff. Locations and dates of their capture in <br />1995 matched those of previous collections of these species lMiller et al. 1982, <br />Wick et al. 1 991), demonstrating that both species use the Little Snake River on a <br />regular basis. We found no evidence of their occurrence in the middle or upper <br />reaches even though a Colorado pikeminnow was previously captured at about <br />