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<br />unregulated snow-melt river in Montana, were consistently less than 5% per day during the <br />baseflow period. <br />Repeated flushing of the varial zone prevents establishment of food webs and resting areas <br />for small fish that are required to support riverine fisheries. Weisberg et al. (1990) 4~monstrated <br />that standing crops of zoobenthos increased 100 fold in one year in a regulated river after <br />eliminating peaking operations at the dam and thereby reducing the devastating ecological effects of <br />unnatural, short-term flushing of the varial zone. Repeated flushing also removes plant growth <br />nutrients and alters the natural thennal insolation of shallow backwaters that are especially <br />important for bioproduction of low velocity food webs in general, and for growth of squawfish and <br />razorback sucker specifically. <br />In spite of the laudable reregulation effort by operators of Flaming Gorge Dam to stay <br />within flow windows (Figure 14) determined to maximize areas of backwater habitats in the alluvial <br />nursery areas of the Green River during summer and fall, 1992, peaking operations still caused <br />considerable diel fluctuation of river stage (e.g., Figures 12-14). I infer that backwaters thought to <br />be protected by these flow windows were in fact flushed or, at least, dramatically fluctuated <br />repeatedly during the late summer, 1992. Data presented by Graboski and Heibert (1989) suggest <br />that the food webs in the backwater environments of the Green River are not very productive. As <br />noted above, these backwaters should contain rooted aquatic plants and a biodiverse, productive <br />invertebrate and fish food web. I realize that the fluctuations shown in Figures 12-14 are <br />considerably reduced from operations in the past. Nonetheless, development of stable, productive <br />food webs in the backwaters probably have not occurred as a consequence of reregulation of the <br />Flaming Gorge releases. Moreover, they will not likely ever be very productive, unless flow <br />fluctuations can be eliminated. Empirical information with which to fmnly judge the productivity of <br />backwater food webs as influenced by regulated baseflow regimes throughout the Upper Colorado <br />River Basin is sorely needed and should be approached in the dynamic time and space context <br />described above. <br /> <br />35 <br /> <br />'. <br /> <br />'", <br />