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to the south in Mexico as previously discussed (and Figs. 2,8,12). Also, a 11111 allele at PGM (Fig. 11) is unique to the Colorado Basin among samples studied and further contributes to that distinctness. However, as exemplified by UPGMA analyses (Figs. 13-17), samples from basins to the east, the Guzman and Rio Grande-Pecos, may be genetically more similar to lower Colorado Basin samples, exclusive of the Lake Mohave and Grand Canyon samples, than are those from the upper Colorado Basin. Because this similarity is based on shared alleles irrespective of comparison to presumably more distantly related species, which might shed light on the primitive or derived state of respective alleles, it is not necessarily a reflection of relationships among populations sampled as will be discussed later in the Integrated Summary. Samples from the lower Colorado Basin and the basins eastward share many alleles that also occur in the upper Colorado Basin samples but additional alleles are restricted to the middle and upper portions of the basin at some loci; this is true at AK- 1 (alleles 1121, & 116"), CBP-1 (allele "211), CK-A (allele "2"), and IDHP-2 (allele 11111)(Figs. 2,4,5,8). The regional distinctions within the Colorado Basin as compared to easterly basins apparent in the UPGMA analysis doubtless owe to the presence of these additional alleles. While differences between samples from the lower Colorado Basin and basins to the east are small, the Guzman and Rio Grande-Pecos samples lack the slower I'1', allele at PGM (Fig. 11) which is pervasive throughout the Colorado Basin; a probable fixed difference at PK-1 (Fig. 12, the single score for the 11211 allele in the Aravaipa Cr. sample is regarded as a possible artifact) further separates Colorado Basin samples from Rio Grande-Pecos samples but not the small Guzman sample as previously noted. Lower Colorado Basin WHile samples of Gila from the Gila and Bill Williams systems comprise a considerable amount of allelic variation, they are characterized overall by a lack of fixed differences. This holds despite the amount of attendant morphological variation and relatively isolated nature of populations. Consequently, UPGMA analysis (Figs. 13-17) generally clustered population samples from these basins as well as the Chevelon Creek sample from the Little Colorado system and the Pahranagat sample from the Pluvial White system of Nevada (Fig. 1) to the exclusion of others with the following exceptions. The small Aqua Fria (AF) sample (n=2) clustered outside all the above samples as well as those from the Rio Grande-Pecos, Guzman, and upper Papigochic (Yaqui dr.) samples (e.g., Fig. 14). Removal of this inadequate (and possibly non Gila, see below) sample from the analysis (Fig. 15), 9