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because the attendant effects were so ecologically devastating (cf. Regier et al. 1989). <br />In most, if not all cases, precious little information about the connectivity of these large <br />rivers was recorded before major human disturbances took place. However, a number <br />of carefully researched case histories provide insightful syntheses of the interactive <br />nature of human and natural disturbances on the ecology of river systems (reviewed in <br />Davies and Walker 1986). <br />Rather than attempt to summarize the many important inferences of these and <br />many other studies chronicling human disturbance in catchments, we present below a <br />single case history of a large catchment that retains many pristine attributes but is <br />threatened by a variety of interactive effects. In this case the cumulative effects may not <br />be so profound that an ecosystem level understanding might be very productive in <br />fostering a new management ethic. The goal is to sustain the natural ecological <br />connectivity of the system. We use this example to set the stage for articulation of some <br />new approaches to that goal that may be useful elsewhere. <br />A Case History of Interactive Effects on Ecosystem Connectivity <br />Background <br />The Flathead River-Lake Ecosystem in northwest Montana provides a good <br />example of a tightly coupled system where natural and human disturbances are clearly <br />interactive. Understanding of this catchment is based on ecological studies by <br />scientists at the Flathead Lake Biological Station (a field station of the University of <br />Montana), where biophysical data have been routinely collected since 1896, and a <br />wide variety of management-oriented research has been conducted by Tribal, State <br />and Federal agencies (reviewed by Stanford in press). Salient points are summarized <br />here. <br />This 22,000 km2 catchment is dominated by runoff from the myriad of tributaries <br />that feed the 6th order Flathead River (mean annual discharge = 340 m3 sec-1), which <br />flows through 496 km2 Flathead Lake (Figure 3). Water quality in this river-lake system <br />is extremely good; solute concentrations and bioproduction are uniformly low <br />(oligotrophic), waters are usually highly transparent (secchi disk readings in Flathead <br />Lake routinely exceed 15 m autumn and winter) and native fisheries are healthy. Less <br />than 80,000 people reside in the entire catchment and no major industrial or <br />agricultural sources of pollution currently exist. The Flathead River dominates the <br />inflow of solutes and particulate materials that influence water quality, structure food <br />webs and drive bioproduction in the lake. For example, the river provides 65% of the <br />annual load of bioavailable phosphorus reaching the lake. Six of the 10 native fishes <br />10