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<br />Squawfish Population Viability Analysis --July 1993 <br /> <br />Page 6 <br /> <br />very important demographic variable. It modulates the stochastic variation <br />of gene frequencies (genetic drift). Demography also controls the <br />"memory" of a population. Only with a long-lived species can the <br />population "remember" past environmental events. That is, a good year <br />might produce a very numerous age class that will be recognizable in the <br />age/size structure of the population for years to follow. This is related to <br />the Storage Effect, considered later. <br /> <br />1.5 Different Historical Demographic Regimes <br /> <br />The Colorado squawfish has evolved as a top carnivore in a wild river <br />system. It's natural demography (pre-fishing by Homo sapiens) should <br />reflect these conditions, since this would be major part of its evolutionary <br />history. This natural regime was interrupted a few hundred years ago with <br />the entry of white man into the river basin. Until the last twenty-five <br />years, the Colorado squawfish was subject to an ever increasing harvest <br />pressure. Beginning in the 1930' s, the underlying stream ecology for the <br />Colorado squawfish was seriously altered with the construction of a series <br />of great dams. Almost simultaneously, the food web of the system was <br />widened and complicated by the introduction of exotic fishes that acted as <br />competitors and predators on juveniles, and possibly as prey. Most <br />recently, in 1974, the Colorado squawfish was declared endangered under <br />federal law, which certainly reduced the harvest pressure on the Colorado <br />squawfish. <br />