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RELATIONSHIPS OF BIRDS, LIZARDS, AND NOCT[.JRNAL RODENTS TO THEIR HABITAT IN TUCSON, ARIZONA <br />Statistical Analysis <br />Habitat ijariables. I defined associations <br />between 15 of the habitat variables using <br />Spearman ranked correlation analysis (Zar 1984, <br />Statsoft 1995). 1 was restricted to 15 variables <br />because several of the original variables were <br />intended for descriptive purposes only and several <br />others were not abundant enough to include. All <br />15 variables were retained for ensuing correlation <br />analyses despite potential inter-correlations <br />because I wanted to identify the association each <br />habitat variable had with species abundances. <br />Inter-correlated variables were not allowed to <br />enter predictive models. The alpha level for this <br />and all statistical tests was set at P <_0.05. <br />Breeding Birds. Bird species and community <br />variables generated at each census plot were: <br />abundance of each species (averaged over the 4 <br />visits), total abundance, and species richness for <br />each of the following 3 groups of resident <br />breeding birds: <br />1. Non-native species; <br />2. Native species; and, <br />3. A guild comprised of birds that were <br />insectivorous, tree or shrub foliage gleaners, <br />and shrub nesters. <br />I selected feeding and nesting substrates as <br />guild delineators to remove the subjectivity in <br />guild selection that has been a major criticism of <br />guild usage (see Holmes et al. 1979, Jaksic Y981, <br />Johnson 1981, Severinghaus 1981, Verner 1984). <br />I determined habitat associations for birds by <br />Gamma correlation analysis of abundances of <br />individual species with habitat variables. Fifteen <br />ranked habitat descriptors were included with <br />ranked abundances of 21 species of birds which <br />met sample size requirements. I chose Gamma <br />correlations because there were many ties in the <br />bird abundance data, and this statistic takes ties <br />into account, whereas Spearman and Kendall Tau <br />coefficients do not (Statsoft 1995). Significant <br />correlations with coefficients >_ 0.5 were retained. <br />I then identified the relationship between each <br />habitat variable and the bird species it correlated <br />with by graphing mean bird species abundance <br />against percent (or distance) classes of each habitat <br />variable. I created classes that were meaningful to <br />land managers and which evenly distributed data <br />among groups. <br />I used correlation and forward stepwise <br />multiple regression to develop predictive habitat <br />models for the 3 bird groups (Statsoft 1995). I <br />used Spearman ranked correlation to determine <br />associations between the bird community <br />descriptors and habitat variables. Habitat <br />variables that were correlated with species richness <br />of each of the 3 bird groups were subjected to <br />stepwise multiple regression analysis. <br />I examined 4 residual diagnostics for each <br />regression analysis. I plotted and removed <br />outliers, which were defined as those cases whose <br />standardized residuals were > 2 standard <br />deviations from the mean residual value. Second, <br />I evaluated Cook's distance (Cook 1977) for each <br />case. Cook's distance is a measure of the affect a <br />case has on the value of the regression coefficient <br />and should be roughly equal for all cases. Third, I <br />examined the Durbin-Watson statistic (Durbin and <br />Watson 1951), which identifies whether cases are <br />independent by analyzing the degree of <br />correlation between adjacent residuals. Last, I <br />reviewed both normal probability plots of <br />residuals and plots of predicted versus residual <br />scores. These indicate if residuals are normally <br />distributed, and test the assumption of a linear <br />relationship between the independent and <br />dependent variables, respectively. <br />I verified the predictive ability of each <br />regression equation by cross validation (Meter et <br />al. 1990). I randomly split the original data into 2 <br />subsets. The equation for the predictive model <br />was generated using 80% of the data, then this <br />model was used to predict values for each case in <br />the 20% subset. I determined the mean squared <br />prediction error (MSPR) and compared it to the <br />error mean square (MSE) of the original data <br />subset. The closeness of these values indicated the <br />extent to which the MSE was biased and gave an <br />indication of how accurately the model should <br />predict the value of the response variable using <br />new data. <br />In addition, I assessed the similarity of slopes <br />in the 2 data subsets by comparing the residual <br />scores between them using an independent <br />samples t-test. If the slope of the second data <br />subset differed from the first, then the scatter of <br />data points would have had a different <br />arrangement than that of the first; i.e., the residual <br />values of the second subset would deviate more <br />from the predicted slope than the residual values <br />for the model building data set. <br />L O ARIZONA GAME Fi FISH DEPARTMENT, TECH. REP. 20 SZEPHEN S. GERMAINE 1995 <br />