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<br />interrelationships between the organisms which process the <br />coarse particles (called shredders) and the microbial <br />flora which colonize the particles have been demonstrated <br />(Cummins, 1973). This processing rate (by structural <br />components) is a quantifiable behavioral characteristic of <br />any river system which is dependent upon temperature, <br />turbulence, pH, water hardness and ionic constituents <br />(Lush and Hynes, 1973). <br /> <br />The organisms which feed on the FPOM produced are <br />called collectors. They aggregate the small particles by <br />their feeding activities. A preliminary analysis of the <br />macroinvertebrate data collected on the White River <br />indicates that the dominant types of invertebrates are <br />collectors dependent upon the heterotrophic food chain <br />(Figure 9). <br /> <br /> <br />FILTER FEEDERS <br /> <br />r~'\ <br /> <br />I '\-~ DETRITlVORES <br />.... I \ !\ ~ATORS <br />~ \ / , <br />. 'A~ I " r\ ..~ <br />~~ HERBIVORES ,'\, '~/ <br />" '\ ~.... <br />\~ I / '~ <br />, .. <br />.. <br /> <br />Figure 9. The distribution (% composition) of the major <br />functional groups of macroinvertebrates in a riffle habitat <br />at Southam Canyon in the White River during 1981. <br /> <br />Respiration rates and absence of invertebrates from <br />decomposing leafpaks indicate microorganisms are very <br />important in the regeneration of this allochthonous <br />detritus for nutrients (Figure 8). <br /> <br />The impact of physical-chemical factors on the <br />distribution of macroinvertebrates is usually indirect, <br />with the major controlling factor being the quantity or <br />quality of food. In general, invertebrates are adapted to <br /> <br />277 <br />