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<br />278 <br /> <br />Y.K. CONVERSE ET AL. <br /> <br />12 <br />10 <br />8 <br />6 <br />4 <br />2 <br />o <br />0.6 <br />0.5 <br />0.4 <br />0.3 <br />0.2 <br />0.1 <br />0.0 <br />10 <br />80 Cover (%) <br /> <br />Depth (m) <br /> <br />o <br />8 00 <br />R oi~ <br />~ el odf> o:i.9 <br /> <br />0(1 <br />:: 0 ~ rID <br />o80QoO <br /> <br />00 <br /> <br />L v <br /> <br />I <br />Velocity (m/s) <br /> <br />o 00 <br /> <br />~O') 0 <br />o~og 0 'cPo <) <br />e o~ o~ 0 0 , _ <br />o 0 0 Q) 0 > (J <br />o 8, 'OUR 0 <br /> <br />60 <br />40 <br />20 <br />06 <br /> <br />00 0 0 <br />o 00 <br />o 0 0 <br />o 0 0 0 0 B& 0 00 <br />~oo_ '"' <br />o oOno~o ~ <br />o 0 ~ (]II 00 0 0 <br />0(0 0 00 aD 00 <br />. . , <br />8 10 12 14 16 18 20 <br />Discharge (cms x 1,000) <br /> <br />Figure 6. Overall change in depth, velocity and cover with discharge. All shoreline types are included in each graph <br /> <br />DISCUSSION <br /> <br />Relationships between humpback chub and geomorphology <br /> <br />Because of the strong influence of geology on the Colorado River ecosystem and the endemic nature of <br />the native fish community, relationships between aquatic habitat and river geomorphology must be <br />understood to manage in a manner which is sympathetic for the welfare of resident fish populations. <br />These relationships seem particularly important to the humpback chub because of its apparent strong <br />evolutionary and ecological ties to geomorphic structure at different life-history stages and different <br />spatial scales (Valdez and Clemmer, 1982; Valdez and Ryel, 1995). In a different region, Rabeni and <br />Jacobson (1993) found a similar relationship. They determined that centrarchid distribution in Ozark <br />streams was influenced by geomorphology on a reach scale and velocity, depth and substrate at a local <br />scale. <br />In this study, we showed that reaches and shoreline types differed physically and that subadult <br />humpback chub presence and abundance in Grand Canyon were related to these geomorphic differences, <br />particularly among shoreline types. This study also demonstrated that subadult humpback chub were <br />specifically associated with a high frequency of cover in channel margins. In fact, the pattern of fish <br />distribution and abundance among shorelines parallels the frequency of cover among shorelines, with <br />vegetation, talus and debris fan shorelines having more cover and greater fish densities than bedrock, <br />cobble and sand shorelines. <br /> <br />(eJ 1998 John Wiley & Sons. Ltd. <br /> <br />Re~ul. Rivers: Res. M~mt. 14: 267-284 (1998) <br />