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<br />INTRODUCTION <br /> <br />The Colorado River drainage is naturally-inhabited by a <br /> <br />comple:< <br /> <br />of cyprinids within the genus GiJ.a. <br /> <br />Currently~ three <br /> <br />species are recognized on the basis of external <br /> <br />morpholoqy: <br /> <br />the <br /> <br />humpback chub~ <br /> <br />cypha Miller; the bony tail chub~ ~. eleQans <br /> <br />Q. <br /> <br />Baird and Girard; and the roundtail <br /> <br />chub~ ~. robusta Baird and <br /> <br />Girard. <br /> <br />Changes in water quality have so inhibited reproductive <br /> <br />success <br /> <br />(Valdez <br /> <br />and Clemmer~ <br /> <br />that the bony tail <br /> <br />1982)~ <br /> <br />humpback chubs are now protected under the Endangered Species <br /> <br />Act. <br /> <br />Competition from introduced fishes may also have influenced <br /> <br />population declines <br /> <br />(Holden and Stalnaker~ 1975; Tyus et al., <br /> <br />1982) . <br /> <br /> , <br />We have little understanding of the phylogenetic relation- <br />ships among the Colorado River Gila. The three recogni~ed <br /> ! <br /> i <br />species comprise closely-related (Uyeno, 1960; Holden i <br />a group ~nd <br /> <br />Stalnaker~ <br /> <br />species status is not clear. <br /> <br />1970) , <br /> <br />but biological <br /> <br />The Colorado River Gila complex <br /> <br />is notable for its varia~le <br /> <br />morphology in relation to ecology (Miller, 1946) and intergr~de <br /> <br />forms are numerous (Holden, <br /> <br />1968; Holden and Stalnaker, 1970>. <br /> <br />In order to understand species relationships with respect to <br /> <br />this morphological variability as well as to provide a foundat on <br /> <br />for sound management~ certain questions must be addressed: <br /> <br />1. <br /> <br />Are the three described s~ecies also true biologi~al <br /> <br />onE'S':' <br /> <br />2. Are species hybridizing <br />are speC1es so variable as <br />SE'('n in Natur-.er:> <br /> <br />1n the wild or, alternate~y, <br />to give the range of body fo~ms <br /> <br />and <br /> <br />i <br /> <br /> <br />~. <br />t <br />