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area did increase to that point. Flows in excess of 9000 cfs rarely occurred during the July and August <br />nursery period, but should be avoided, if possible, to maximize native roundtail chub recruitment. <br />Westwater Canvon <br />A greater diversity of low velocity habitat types was found and sampled in Westwater Canyon <br />than upstream. At the presence/absence level of detection, a multinomial analysis indicated YOY chubs <br />utilized habitats as they were available, i.e., no selection. A closer look at the densities of YOY chubs by <br />habitat type (Figure 9) reveals some habitats were preferred. Interestingly, the density of YOY chubs in <br />backwaters was statistically less than expected. Embayments, characterized as pocket water areas or <br />concavities in the shoreline geology, held greater densities of YOY chubs than was predicted. A greater <br />percentage of effort was expended sampling shoreline habitats in the canyon than upstream - a habitat <br />that was utilized by YOY chubs as it was available. <br />Backwaters were negatively (not significantly) correlated with sample flows. Embayments were f <br />available at a wide range of flows and were positively correlated with flows at the time of sampling. <br />Although very few larval chubs were collected throughout this study, it is likely these embayments or <br />shoreline pocket waters that provided habitat for the very early life stages of native Gila spp. during the <br />high flows were associated with the descending limb of the hydrograph. Embayment habitat availability <br />was a function of instantaneous flow /river stage. These habitats are not formed by sediment transport, <br />but rather a simple process of the river inundating or stranding shoreline geology. <br />One of the objectives of this study called for a description of the relationship between <br />geomorphological processes of sediment transport and nursery habitat formation and maintenance in <br />Westwater Canyon. We did not coordinate with a geomorphologist in this study. UDWR did contract <br />with geomorphologists to describe the relationship of low velocity habitat (eddys) and flow in Desolation <br />/ Gray Canyon where numerous backwaters form in eddy return channels as flows drop in the summer <br />(Orchard and Schmidt 1998). The short stretch of Colorado River flowing through Westwater Canyon <br />characterized by steep gradient and confined channel configuration, is not an area of sediment <br />deposition. The availability of backwater habitat was negatively correlated with the magnitude of the <br />spring peak, but was more likely a function of late summer flows. It was our observation that the <br />magnitude of the spring flood had very little effect on the availability of low velocity habitat in general <br />in Westwater Canyon. Backwaters and other habitats associated with sand bars were sufficiently scarce <br />that it quickly became apparent that humpback chub could not depend on them greatly as nursery areas. <br />A multinomial analysis of habitat selection by young chubs indicated they were not used as much as was <br />statistically expected. [If future studies indicate humpback chub spawn on cobble bars in Westwater <br />Canyon, a geomorphological perspective of how those habitats are maintained could be warranted.] <br />Our observations were that YOY chubs in the canyon used whatever low velocity habitat was <br />available. In a narrow canyon like Westwater, where relatively small changes in flow result in large <br />changes in stage, specific habitat stability is poor. If larval and YOY chubs are going to persist in <br />Westwater Canyon, and they do, they need to be opportunistic in their habitat use. Day and Crosby <br />(1997) also found that YOY chubs did not select habitat types preferentially in Deso/Gray Canyon. <br />Arizona Game and Fish Department (1996) YOY surveys in the Grand Canyon found higher percentages <br />(not higher densities) of humpback chub in the beachfront species assemblages (similar to our shoreline <br />designation) than in backwaters. This opportunistic use of low velocity areas is likely a life history <br />strategy more common of G. cypha than G. robusta. High densities of YOY roundtail chubs were found <br />in backwater habitats upstream of Westwater Canyon. The relationship between the availability of low <br />velocity habitats and summer flows in Westwater Canyon reported herein, indicates that roundtail chub <br />(and non-native cyprinids) likely do better during the low flow years. This finding, in part, describes the <br />mechanism of reproductive isolation for these two native chubs, which coincides with the trends in the <br />long term monitoring data sets. <br />23 <br />