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is at the top of the15-mile reach (Fig. 6). <br />Thus, in terms of forage and suitable temper- <br />ature regime, the 15-mile reach may be <br />located in or near the most optimal part of <br />the Colorado River for Colorado pikemin- <br />now. <br />Low water temperature might also in- <br />hibit successful spawning by pikeminnow in <br />reaches upstream of Palisade. In De Beque <br />Canyon, temperatures necessary for spawn- <br />ing are met, but the overall thermal regime <br />may be less than optimal in some years. The <br />initiation of spawning in the Colorado River <br />generally occurs when river temperatures <br />exceed 18 ° C and spawning activity lasts 3-8 <br />weeks (Trammell and Chart 1999). Hamman <br />(1981) reported that females held in race- <br />ways did not spawn until temperatures ex- <br />ceeded 20°C. In controlled experiments, <br />Marsh (1985) found that for eggs spawned <br />25 <br />U <br />Ch <br />20 <br />N <br />L <br />I? <br />a? 15 <br />L <br />CL 10 <br />E <br />a? <br />5 <br />and fertilized at 18 ° C, hatching success and <br />embryo survival was highest at 20°C. <br />Although some spawning occurs at cooler <br />temperatures (see Bestgen et al. 1998), <br />most spawning in the Colorado, Green, and <br />Yampa rivers occurs at water temperatures <br />between 18-22'C (McAda and Kaeding <br />1991, Tyus 1990, Bestgen et al. 1998, <br />Anderson 1999, Trammel and Chart 1999). <br />Analyses of temperature records at Cameo <br />(1986-1996) indicate that 18°C is exceeded <br />in both wet and dry years, but in wet years, <br />20'C may be only briefly attained (Fig. 7). <br />Thus, in some years suitable conditions for <br />spawning may be of short duration. In <br />addition, the delayed annual warming of <br />upstream reaches will likely delay the onset <br />of spawning, resulting in larvae having a <br />shorter first-year growing season than if <br />produced in more downstream reaches. <br />20 C Y STATE LINE <br />SPAWNING 18 C ---- - _?,X__ _ <br />CAMEO <br />1993 <br />MAY JUN JUL AUG SEP OCT <br />Month <br />Figure 7. Main channel temperatures (May-October) at two sites in the Colorado River during <br />a wet year, 1993. Measurements are from USGS gauging stations at Cameo and the <br />Colorado-Utah state line. Plotted daily values are the average of the daily maximum and <br />minimum readings. Timing and duration of temperatures in excess of 18 ° and 20 ° C (see text) <br />at the two sites are shown for comparison. <br />11