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70 <br />60 <br />Q 50 <br />c <br />40 <br />30 <br />20 <br />133 182 200 230 93 117 <br />River Mile <br />Figure 6. Mean annual thermal units (ATU) available for Colorado pikeminnow growth at <br />four locations on the Colorado River and two on the Yampa River for comparison. Dashed <br />horizontal lines indicate sites on each river with similar thermal regimes. River mile (rm) 133 <br />is near the Colorado/Utah state line; rm 182 is 3 miles downstream of the top of the 15-mile <br />reach; rm 200 is 7 mules upstream of Cameo; rm 230 is at Rulison. On the Yampa River, rm <br />93 is at Juniper Springs (the Yampa reach where densities of Colorado pikemmnow are <br />higghest); rm 117 is near Sheep Creek (the site of Nesler's [1995] most upstream Colorado <br />pikeminnow capture). Data are from Osmundson (1999a). <br />only 67% of that at the state line, and at <br />Rulison, only 52% (Fig. 6). <br />Although abundant forage may compen- <br />sate for slow growth caused by low tempera- <br />tures, there is a point at which temperatures <br />are so limited that additional food cannot <br />make up for the temperature shortfall and <br />growth slows. What defines the upstream <br />range of this species is likely a temperature <br />threshold, below which annual growth is <br />insufficient. Colorado pikeminnow can likely <br />gauge about where this minimum thermal <br />regime occurs and individual home ranges <br />are therefore established no further upstream <br />of this point. <br />In the Yampa River, fish have unre- <br />stricted upstream access. Nester (1995) <br />found adult Colorado pikeminnow in the <br />Yampa as far upstream as river mile (rm) <br />117. This location has an average ATU <br />value of about 41. On the Colorado River, <br />the location with the same mean ATU <br />occurs at approximately rm 199, or about <br />14 miles upstream of Palisade. Using the <br />Yampa River as a model, Osmundson <br />(1999a) suggested that Colorado pikemin- <br />now in the upper Colorado River should be <br />able to establish home ranges upstream <br />almost as far as De Beque, Colorado, once <br />passage is provided. Also, the thermal <br />regime in the reach where Colorado pike- <br />minnow were most abundant in the Yampa <br />River averaged 46.5 ATU (rm 93). The <br />analogue to this site in the Colorado River <br />10