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6 <br />The catfishes, Ictaluridae, with two species in two genera, <br />present another level of reproductive behavior. The male guards <br />and fans the eggs, and he shepherds the baby fish. Sometimes the <br />female remains and assists the male in caring for the young, <br />producing a monogamous pair. <br />Best known for monogamy and biparental care, however, is the <br />family Cichlidae. Interestingly, the common cichlid in Cuatro <br />Cienegas may prove to be a maternal caretaking species, an <br />unusual state of affairs for fishes in general. This is one of <br />the most ubiquitous fishes in the basin, and its mating system <br />may well differ among the springs. <br />The Poecillidae is treated last because of its advanced care <br />of young. The female carries them in her ovary where they develop <br />out of harm's way. The mating system is not known in any detail, <br />but fishes in this family are polygynous. <br />Thus we see that the situation at Cuatro Cienegas offers <br />unparalleled opportunities. It is a complex of natural <br />experiments such that the judicious investigator can pick and <br />choose springs appropriate to the hypothesis in question. <br />Because of the shallow clear waters, benign experiments can then <br />be carried out to refine the study. <br />Now I would like to digress to address briefly two issues. <br />One is the re-establishment of populations where they once <br />existed but have become extinct. The other is the issue of <br />extinction in the larger context. <br />According to Sewell Wright it is virtually axiomatic that <br />evolutionary change is hastened by small population size. When a <br />small founder population is introduced into a spring, as has