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This should be of concern, in that it also demonstrates #hat hybrid individuals can be <br />fertile, and that introgressed genes can be perpetuated in the population. Interestingly, <br />only large (i.e., old) native suckers were found in the Little Yampa Canyon reach, <br />whereas a greater size distribution (and thus age structure) was maintained in the <br />introduced White Sucker (and to some extent hybrids). Lower reaches of the Yampa <br />River, where natives are more abundant, were not evaluated for this trend. However, <br />the hybrid zone appears localized and its effects dwindle as one moves downstream. <br />Thus, the spread of individuals with alien genomes may not be of immediate concern. <br />INTRODUCTI©N <br />Numerous factors have contributed to the decline of indigenous fishes in western <br />North America (Miller, 1961; reviewed by Hinckley and Douglas, 1991). Yet, the most <br />serious stem from planned or impromptu introduction of alien species (both vertebrate <br />and invertebrate). There are 36 indigenous fish species that comprise the Colorado <br />River Basin ichthyofauna, and they have been subjected to over 70 nonnative fish <br />introductions (Harsh and Douglas, 1997). Three indigenous species are now extinct, <br />and 22 are either federally endangered or threatened. As a result, southwestern United <br />States is now recognized as a major "hotspot" for reduction of endemic fish biodiversity <br />(Dobson et al., 1997:figure 1 c). <br />There are several mechanisms by which exotic fishes impact indigenous ones. <br />From an ecological standpoint, overt predation of the former upon the latter is most <br />visible and apparent (Heffe, 1985; Marsh and Langhors#, 198$; Marsh and Douglas, <br />1997). Here, impacts are immediate and severe, such that native fish densities become <br />depressed and recruitment curtailed (Hinckley, 1973; Bestgen and Propst, 1989). <br />However, exotic fishes also affect endemics in ways that occur more slowly and over <br />longer periods of time, yet with results equally as devastating as overt predation. <br />Exotics, for example, may out-compete and ultimately replace endemics, particularly <br />when the realized niches of the two overlap considerably (reviewed by Ross, 1991; <br />4 <br />