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condensed into a capture history (i.e., CH) matrix (Burnham et <br />al., 1987; Lebreton et al., 1991). Fifty-seven matrices were <br />derived (three camps over 19 trips). <br />Closed population estimates.---Population estimates were <br />generated from each CH-matrix under assumption that the three <br />stream reaches contained closed populations. This was appropriate <br />given the brief sampling period at each camp (see Otis et al., <br />1978), and because only adults were censused. Closure was tested <br />by examining numbers of individuals tagged within one reach then <br />recaptured within a second reach during the same trip. Nine <br />different closed-population estimates were derived from each CH- <br />matrix using an updated (30 Dec. 1991) version of the computer <br />program CAPTURE (G. C. White, D. R. Anderson, K. P. Burnham, and <br />D. L. Otis, Los Alamos Natl. Lab., 1982, unpubl.). Models and <br />assumptions are explained in Otis et al. (1978), Pollock et al. <br />(1990), and Nichols (1992). The single best-fitting population <br />model, as indicated by goodness-of-fit tests and comparisons <br />between competing models, was retained. In this first analysis, <br />population estimates were made relative to one another by <br />dividing each by length of reach (in km). ANCOVA (Proc GLM; SAS, <br />1985) then contrasted relative population estimates by reach, <br />using fishing effort as a covariate. <br />In a second analysis, tag/recaptures were evaluated for the <br />entire LCR (rather than by reach). Here, 19 CH-matrices were <br />generated, one for each month of study. Again, the single best- <br />fitting population model was retained. ANOVA was used to test the <br />19 estimates against those summed by reach for each month. The <br />hypothesis under test is that monthly estimates are not <br />significantly different from those summed by month over reaches. <br />Finally, a third analysis collapsed all tag/recaptures into <br />a single CH-matrix (i.e., each column of the CH-matrix <br />represented a single month). Here, five best-fitting estimates <br />were retained. However, assumptions of closure may be violated in <br />this analysis by movements of G. cypha into/from the mainstem <br />Colorado River over the 19-month study interval, and by <br />recruitment of juvenile chubs into the adult population. Thus, <br />while this analysis is a logical culmination of population <br />estimates (a) by reach, (b) by month summed over reach, and (c) <br />solely by month, results are heuristic rather than practical. <br />RESULTS <br />Fishing effort and unadjusted population estimates.---Fishing <br />effort differed significantly among reaches (F=6.40; P < 0.0035; <br />Proc GLM; SAS, 1985), with effort at Salt Canyon greater than <br />that at Confluence (Sidak's multiple range test; SAS, 1985). <br />Efforts at Salt and Powell Canyon reaches were statistically <br />similar, however. Population estimates (normalized by river km) <br />4