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will not be possible. Wydoski and Wick (1996) discussed in depth the ecological <br />importance of floodplains to riverine fish communities and identified strategies <br />to enhance or restore flooded bottomland habitats to assist recovery of <br />endangered fishes in the Upper Colorado River Basin. They felt that it is <br />imperative to integrate the control of nonnative fishes and <br />augmentation/restoration stocking of captive-reared razorback sucker with habitat <br />enhancement/restoration efforts. Enhancement or restoration of biological, <br />chemical, and physical conditions that are perceived to be important in nursery <br />sites for razorback sucker can be accomplished by: 1) removal of levees to allow <br />periodic flooding of lowlands, 2) providing a longer duration of inundation in <br />floodplains, embayments, and backwaters by reoperation of releases from <br />reservoirs, and 3) reconnection of the main channel with off-channel oxbow lakes <br />and ponds. The removal of levees coupled with higher spring releases that mimic <br />the historic hydrograph from upstream dams provided the best way to reconnect <br />mainstem and tributary rivers with productive floodplain areas. Such removal <br />should be done on low fl oodpl ai n terraces on public properties that can be easily <br />reconnected with the main channel and where thorough, followup evaluations can <br />be made. <br />However, some nonnati ve fi shes that are establ i shed i n the upper basi n wi 11 <br />flourish in productive off-channel habitats. Tyus et al. (1982) estimated that <br />76.4% of the fish community in the Upper Colorado River Basin is composed of <br />nonnative fishes. Predation and competition on larval razorback sucker by <br />nonnative fish is probably the primary factor that drastically reduces or <br />prevents recruitment of this species today. If the duration of inundation of <br />off-channel habitats is short, razorback suckers will not grow to a size where <br />predation and competition by nonnative fishes is no longer a principal factor in <br />their survival. Floodplain habitats that are ephemeral will not provide enough <br />time for razorback sucker larvae to reach a length of 1 inch. Since nonnative <br />fish species dominate backwater habitats (up to 90-99% based on ISMP data [McAda <br />et al. 1994]), razorback sucker larvae that enter the river as ephemeral <br />floodplains recede will be extremely vulnerable to predation since razorback <br />sucker larvae constitute the only wiggling food items in the spring. Based on <br />Papoulias and Minckley's work (1990), about 2 months is required for razorback <br />suckers to reach a length of 1 inch. Razorback sucker that are about 1-inch long <br />are no longer vulnerable to predation by some predators such as red shiners based <br />on the gape size work by Todd Crowl. If the duration of flooding these habitats <br />is longer, nonnative fishes will flourish and continue to adversely affect larval <br />and juvenile razorback sucker. Wydoski and Wick (1996) agreed with Stanford <br />(1994) who believed that the negative nonnative fish interactions may limit the <br />recovery of razorback sucker regardless of possible streamflow provisions. They <br />also felt that recovery of this species may be limited unless adequate control <br />measures can be implemented in upstream, and particularly in downstream reaches <br />of main channels, where razorback sucker larvae drift as the flooded areas recede <br />following high streamflows. If adequate control of nonnative fishes cannot be <br />achieved, they suggest that human intervention may be required to rear razorback <br />suckers in predator-free, off-channel habitats so that their populations can be <br />maintained through augmentation stocking. <br />Of the four "large-river" endemic, warmwater fishes that are listed <br />endangered in the Upper Colorado River Basin, the Recovery Program considers the <br />razorback sucker the highest priority species for recovery. While razorback <br />23