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are additional measurements that could be made that may provide a better picture of <br />interstitial void space. The barrel sampler (Milhous et al. 1995) reliably samples surface and <br />subsurface materials in gravel- and cobble-bed rivers. Also, bulk density can be used to <br />monitor relative changes in void space: an increase in weight of a bed sample indicates loss of <br />voids. Similarly, water displacement can be used to measure void volume in river bed <br />samples. These techniques are relatively time consuming; however, limited use of one or <br />more of these methods concurrent with depth-to-embeddedness sampling may provide a <br />predictive relationship between depth-to-embeddedness and void volume. <br />We did not encounter sandy areas between rocks of the surface layer as one might encounter <br />in areas of high embeddedness. Thus, our Wolman pebble counts used to describe the <br />particle size distribution of the surface layer were able to deal exclusively with particles > 8- <br />10 mm in diameter. If future monitoring reveals high intrusion of fines between rocks of the <br />surface layer, some additional technique will need to be applied to provide some measure of <br />the area of silt or sand between rocks. <br />As previously mentioned, a March date should be added to the annual sampling regime. This <br />would allow determination of sedimentation over winter and would better isolate the time of <br />greatest flushing during runoff. <br />To better define the relationship between embeddedness and food availability for fish species <br />of concern, it is recommended that appropriate biological sampling be done concurrently with <br />embeddedness monitoring. Time and funding constraints did not allow such sampling during <br />the current study. This sampling should be designed to address the following question: <br />How does the productivity of key invertebrate species (those used as food by target fish) <br />respond to changes in substrate embeddedness? Combined with fish food habit data, a <br />physical process-biological response model could be developed for fish carrying capacity in <br />much the same way that the Harvey et al. (1993) model links flows and geology to Colorado <br />squawfish spawning site selection. Knowledge of the carrying capacity of various reaches is <br />the missing link in determining realistic recovery goals for the endangered fish. <br />We recommend that: <br />1) monitoring be continued for the next several years using the protocol described here, <br />2) an additional sampling date during March or early April be included in the sampling <br />design, and <br />3) monitoring of mean standing stock biomass of key invertebrates (determined from fish <br />food habit studies) be conducted concurrently with embeddedness monitoring. <br />29