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Disturbance can be an important determinant of the foodweb that develops in the benthos. <br />Wooton et al. (1996) found that predator-resistant ca.ddisflies (Dicosmoecusgilvipes) greatly <br />increased in abundance in northern California rivers after flood frequencies were reduced <br />from river regulation; this resulted in diminished energy in the food chain flowing to <br />predatory fish by diverting most of the energy to predator-resistant grazers. The end result <br />was lowered food availability for juvenile steelhead (Onchorynchus mykiss). Thus, reduction <br />in invertebrate numbers or overall invertebrate productivity is only the most obvious means <br />by which native fish faunas may be affected by changes in flow regime -- changes in <br />invertebrate species composition may be equally important. <br />While high spring flows are still the norm in the upper Colorado River, even with flow <br />regulation, such regulation has resulted in a reduced frequency of years with flows of <br />sufficient magnitude and duration to perform the dual role of cleaning the bed of fines and <br />disturbing the benthic community. To what extent this reduction in frequency ultimately <br />affects food availability for native fish is as yet unknown. We suspect that periods of back- <br />to-back low-flow years would allow for the greatest changes to potentially occur. Studies <br />are needed that will increase our understanding of the relationship between flows and the <br />dynamics of invertebrate communities and foodweb linkages. Our study of embeddedness in <br />relation to flows is only a first step in developing such an understanding. <br />CONCLUSIONS <br />1) The sampling regime designed for and used in this study proved effective at identifying <br />differences in embeddedness within different sites and habitat types and through time. <br />2) Though statistically significant, differences in depth-to-embeddedness detected between <br />strata, bank position, and year were slight during the 1996 and 1997 study periods. <br />3) In general, depth-to-embeddedness was higher in riffles than in runs. <br />4) The years of our sampling represent a period of time when both reaches were freshly <br />cleaned of fine sediments and should therefore provide a good baseline for future <br />monitoring efforts. <br />RECOMMENDATIONS <br />We suspect that 5-6 years of consecutive monitoring will need to be done before trends in <br />embeddedness can be adequately evaluated; the need is for a series of years containing a wide <br />range of flow conditions. <br />Depth-to-embeddedness was used in this study because many samples could be processed <br />relatively quickly by a small field crew and no laboratory time was required. However, there <br />28