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<br />agricultural land. Only a small percentage of the riverbank is stabilized by shrubby vegetation and bank <br />erosion is common. The Maybell gage is located near the upper edge of this stratum at RM 85.8. <br /> <br />Stratum 8 extends from RM 91.6 to RM 135 at the town of Craig, Colorado. In this stratum the valley <br />tends to be more confined than in Stratum 6. Between RM 91.6 and RM 105.0 the valley is wide enough <br />for hay fields and pastures adjacent to the river. The river reach between RM 105 and RM 126 is the <br />Bureau of Land Management (BLM) Little Yampa Canyon Management Unit. Most of the river bottom <br />in this reach is owned by the BLM. The canyon walls are not as confined as Yampa Canyon and a flood <br />plain is typically associated with at least one side of the river. <br /> <br />Description ofYampa River Biotic System <br />The influence of seasonal changes in hydrology on movement and reproduction of large river fishes in the <br />Upper Colorado River Basin is well documented (e.g. Tyus and Karp 1989, Stanford 1994). Hydrology is <br />important in initiating movement to spawning areas (Nesler et al. 1988, Tyus 1990, Karp and Tyus 1990, <br />Modde and Irving 1998) and determining both the formation (Ligon et at 1995, Rakowski 1997) and <br />availability (Orchard and Schmidt 1998) of habitat for fishes in large rivers. Much of our knowledge of <br />hydrological effects on fish in the Colorado River Basin are linked to responses from high and receding <br />flows on fish and habitat (Stanford 1994). Relatively little is known about the importance oflow flows to <br />the maintenance of native fishes in southwestern rivers. <br /> <br />Several studies have shown that native fishes tolerate higher flow events than nonnative fishes in arid <br />streams and rivers in the southwest (Meffe and Minckley 1987, Minckley and Meffe 1987, Deacon 1988, <br />Hoffnagle et al. In review, Muth and Nesler 1993). Hawkins and Nesler (1991) reported that nonnative <br />fishes declined following years with high spring discharge in the Upper Colorado River Basin. In <br />general, fishes native to the southwest have shown a greater tolerance to environmental variabil~ty. Given <br />that native fish tolerate a more highly variable flow regime, seasonal as well as annual variability was an <br />important element in the modified flow recommendation for the Yampa River by Modde and Smith <br />(1995). Poff et al. (1997) suggested that the natural flow regime of all rivers is inherently variable and <br />that this variability is critical to ecosystem function and native biodiversity. Studies in Midwestern (Poff <br />and Allan 1995), Californian (Baltz and Moyle 1993), and Australian streams (Closs and Lake 1996) have <br />suggested that maintaining natural flows regimes, including baseflow variation, is important in <br />maintaining native fish assemblages. This rationale suggests that if environmental variation is reduced, <br />dominant nonnative species could displace or eliminate tolerant, less competitive native species. Because <br />abiotic factors are more influential on ecological processes in highly variable systems such as the Yampa <br /> <br />26 <br />