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<br />associated with foraging behavior. During the spring and summer of 1995, 12 adult <br />razorback sucker were collected in a wetland on the Ouray National Wildlife Refuge <br />(Modde 1996; USFWS unpublished data). Fish presumably accessed the wetland <br />during the peak spring flows when the wetland was connected to the river, and <br />remained in the wetland after the river elevation receded. Razorback sucker have also <br />been observed in floodplain wetlands by Tyus and "Karp (1990) and Bruce Waddell <br />(U.S. Fish and Wildlife Service, unpublished data). In 1993, three fish monitored by <br />telemetry were located at the mouth of a tributary (Ashley Creek) following spawning. <br />Two razorback sucker traveled downstream to the vicinity of the Ouray wetland complex <br />during both years of the study. Historically, floodplain habitat along the Green River <br />between the Yampa and White rivers was inundated during the spring months <br />(approximately 20,000 cfs or 566 m3/s) an average of 2 out of every 3 years (U.S.G.S. <br />records from the gage at Jensen, Utah). As suggested by Tyus and Karp (1990), <br />razorback sucker attraction to wetland outlets and tributary mouths may be related to <br />temperature preference (Bulkley and Pimentel 1983) and productivity (Mabey 1993). <br />Fish implanted with radio transmitters returned upstream to the vicinity of Split Mountain <br />Canyon at the onset of the base flows. <br /> <br />Cueing to natural hydrographic and thermal regimes in riverine environments <br />may effectively concentrate fish at spawning sites, thus increasing the efficiency of <br />successful spawning, particularly within small populations. Razorback sucker in Lake <br />Mohave, Arizona, in the absence of riverine cues, spawned over a longer period of time <br />between November and May (Minckley 1983) with most of the individuals spawning <br />over a shorter period of time. Despite chronological differences, temperature reported <br />for peak razorback spawning in the lower basin, 100 and 150C (Bozek et al. 1984), was <br />somewhat lower but similar for razorback sucker in the upper Colorado River basin <br />(Tyus 1987; Tyus and Karp 1990). Thus, discharge cues are hot needed for <br />reproduction, but flow patterns can initiate movement that may help synchronize <br />spawning adults on historic spawning sites. The latter is important in maximizing <br />reproductive success in small populations. The timing of reproduction is also important <br />if floodplain inundation is necessary to optimize survival of early life stages of razorback <br />sucker (Tyus and Karp 1990; Modde et al. 1996). <br /> <br />Tyus and Karp (1990) identified the Yampa River and Escalante (Jensen) <br />spawning areas, and Tyus (1987) suggested that spawning may also occur near the <br />mouth of the Duchesne River and in Island Park in Dinosaur National Monument. <br />Telemetry data supported the concept that multiple spawning sites existed, including <br />the Yampa River near its confluence with the Green River, the Escalante spawning <br />area, and possibly the lower portion of Island Park. Transmitter-implanted razorback <br />sucker moved to the vicinity of the Duchesne River either prior to or following spawning. <br />However, capture data showed that some razorback sucker occupied the Duchesne <br />River area during the peak spawning period. Tyus and Karp (1990) suggested that <br />razorback sucker showed fidelity to the Escalante spawning area. They observed a <br />razorback sucker, via telemetry, bypass the Escalante spawning area enroute to the <br />Yampa River site. However, fidelity to spawning sites is difficult to ascertain from <br /> <br />22 <br />