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<br />16 <br /> <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />the density of planktonic crustaceans (cladocerans and copepods) averaged about 36 organisms per liter <br /> <br /> <br />during the summer of 1993 (Cooper and Severn 1994a). Cooper and Severn reported a mean of 2 <br /> <br /> <br />organisms per liter for backwater sites and a mean of 36 organisms per liter for open water wetlands. <br /> <br /> <br />Samples of planktonic crustaceans (c1adocerans and copepods) from the Escalante Ranch site on the <br /> <br /> <br />middle Green River, upstream from Jensen, Utah, contained 0 organisms per liter for the main channel, a <br /> <br /> <br />mean of 41 organisms per liter for backwaters, and a mean of 71 organisms per liter for an open water <br /> <br /> <br />wetland (Cooper and Severn 1994b). The mean number of cladocerans and copepods from a backwater <br /> <br /> <br />of the Gunnison River at the Escalante State Wildlife Area, about 5 miles downstream from Delta, Colorado, <br /> <br /> <br />was 11 organisms per liter of water and the mean number from an open water wetland was 25 organisms <br /> <br /> <br />per liter (Cooper and Severn 1994c). Zooplankton (cladocerans and copepods) samples from a bottomland <br /> <br /> <br />(Old Charlie Wash) on the Ouray National Wildlife Refuge contained a mean of 31 organisms per liter of <br /> <br /> <br />water (Cooper and Severn 1994d). Samples taken from the middle Green River and a backwater on the <br /> <br /> <br />refuge did not contain any c1adocerans or copepods. <br /> <br />C. Maior limitinG Factor Affecting EndanGered Colorado River Fish Pooulations. The major limiting factor <br />affecting the populations of endangered Colorado River fishes is the lack of successful recruitment (U.S. Fish <br />and Wildlife Service 1987, 1990a, 1990b, 1991). In the Lower Colorado River Basin, studies have <br />demonstrated that predation by non-native fishes and the lack of food may be limiting the survival of larval <br />and juvenile endangered fishes (Marsh and Langhorst 1988; Papoulias and Minckley 1990, Papoulias and <br />Minckley 1992). Similar conclusions have been reached for the endangered Colorado River fishes in the <br />upper basin (Miller et al. 1982; Maddux et al. 1993). <br /> <br />D. Relation of Food to Larval Fish Survival. Year-class strength of marine fisheries is often related to the <br /> <br /> <br />Match or Mismatch Principle where environmental conditions such as water temperature as well as the <br /> <br /> <br />quality and quantity of food organisms must match the hatching of larval fishes to realize high survival (Hjort <br /> <br /> <br />1914, 1926). In other words, the density, size, time and duration of availability of zooplankton must "match" <br />