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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />The rivers of the Upper Colorado River Basin are turbid and contain large expanses of sand substrate. <br /> <br /> <br />Production of phytoplankton and zooplankton that form the basis for a food pyramid are extremely low in <br /> <br /> <br />the these rivers (Grabowski and Hiebert 1989; Cooper and Severn 1994 a, b, c, and d; Mabey and Shiozawa <br /> <br /> <br />1993). High turbidity obstructs the penetration of sunlight that is needed for phytoplankton production. <br /> <br /> <br />Backwaters and embayments along the main river channels and flooded bottomlands in off-channel areas <br /> <br /> <br />provide favorable conditions for phytoplankton production. Sediments deposited in these areas where the <br /> <br /> <br />water velocity is decreased provide nutrients and sunlight penetrates the clearer water that allows <br /> <br /> <br />phytoplankton to flourish as primary producers and to stimulate production of the food chain. Low velocity <br /> <br /> <br />off-channel habitats become warmer than the riverine environment in the upper basin (Kaeding and <br /> <br /> <br />Osmundson 1988; Osmundson and Kaeding 1989). The combination of nutrients, sunlight penetration of <br /> <br /> <br />the water column, and warmer water temperatures in low velOCity habitats provide the best conditions for <br /> <br /> <br />phytoplankton production in the upper basin. <br /> <br />B. Importance of Low Velocity Habitats to Zooolankton Production. The importance of low velocity habitats <br /> <br /> <br />to the production of zooplankton for fish in riverine environments has not been documented very well. Most <br /> <br /> <br />riverine studies have concentrated on macroinvertebrates occurring in the drift (Waters 1969). Mabey and <br /> <br /> <br />Shiozawa (1993) reported that the most comprehensive studies have been made of the plankton <br /> <br /> <br />communities in the Orinoco River, Venezuela. Mean densitites of c1adocerans and copepods (the most <br /> <br /> <br />abundant taxa) were 421 organisms per liter in the Laguna la Orsinera. Welcomme (1989) summarized <br /> <br /> <br />zooplankton densities in floodplains in a range of 270 to 10,000 organisms per liter. Mabey and Shiozawa <br /> <br /> <br />(1993) documented zooplankton densities in the middle Green River as 0.3 to 1.3 organisms per liter, 1.5 <br /> <br /> <br />to 7.1 in the Ouray backwater, 63.4 at Intersection Wash (another backwater), and 206 to 690 in Old Charlie <br /> <br /> <br />Wash (Woods Bottom) on the Ouray National Wildlife Refuge, located downstream from Vernal, Utah. <br /> <br /> <br />Grabowski and Hiebert (1989) reported 0 to 20 planktonic crustaceans (cladocerans and copepods) per liter <br /> <br /> <br />in the middle Green River channel and 0.02 to 17 organisms per liter in backwaters during 1987 and 1988. <br /> <br /> <br />In an open water habitat of the Moab Slough, immediately upstream on the Colorado River from Moab, Utah, <br /> <br />15 <br />