Research, monitoring reports, and interview com-
<br />ments all suggest that "successful" grass establish-
<br />ment displaces some native plant regeneration. This
<br />was the goal of past range "reseeding" projects-
<br />producinguseful livestock and wildlife forage on land
<br />that would not contain harveetable timber for decades
<br />and would otherwise produce nothing but "weedy"-
<br />and aggressive, persistent grass species were deliber-
<br />ately chosen for seeding (Christ 1934, Evanko 1955,
<br />Friedrich 1947, McClure 1956, Stewart 1973). Sup-
<br />preaeion ofnativeplant regeneration could potentially
<br />reduce browse species for wildlife, reduce watershed
<br />protection in chaparral, and limit the seed bank con-
<br />tributions of annual and short-lived perennial "fire-
<br />followers" in chaparral and Southwestern ecosys-
<br />tems (Conard and others 1995, Keeler-Wolf 1995,
<br />Keeley and others 1981, Loftin and others 1998).
<br />There ie no published research that quantifies the
<br />long-term impacts of poatfire seeding on native plants,
<br />but one monitoring observed that weeping lovegrasa
<br />(Eragrostis curwia) in the Southwest can effectively
<br />suppress native vegetation for years (Loftin and oth-
<br />ers 1998).
<br />In our interviews, forest silviculturiats expressed
<br />major concerns about the impacts of grass seeding on
<br />conifer regeneration. The dilemma between erosion
<br />reduction and conifergrowth is well recognized: "Since
<br />granitica are inherently good tree-growing sites, as
<br />well as being extremely erodible when burned, the
<br />choice between immediate reforestation and long-
<br />termproductivity can be a difficult one" (Van de Water
<br />1998, p. 28). Better understanding of the impacts of
<br />fire and erosion on soil productivity would help ad-
<br />dresathis problem.
<br />Current USDA Forest Service guidelines promote
<br />the use of native species for revegetation projects
<br />wherever practical. Interviewees commented that
<br />native grasses are expensive and not widely available
<br />inthe quantitiesnecessaryfor postfire seeding projects,
<br />and developing seed sources that can provide a range
<br />oflocally adapted genotypes is difficult (Van de Water
<br />1998). In addition, well-adapted native perennial
<br />grasses could provide as much or more competition
<br />with conifers as the non-native species currently in
<br />use. For example, the native Southwestern grass Ari-
<br />zona fescue (Festuca arizonica) greatly reduced the
<br />growth of conifer seedlings (Pearson 1942, Rietveld
<br />1975). One BAER team included the coat of using
<br />herbicide for seeded grass control in BAER calcula-
<br />tions and, as a result, decided against using awell-
<br />adapted native grass and chose to seed cereal barley
<br />(Hordeum wlgare) instead (Griffith 1998). The barley
<br />died out after 1 year except where disturbed by salvage
<br />logging (Griffith 1993; tables 14, 15).
<br />Seeded grasses can benefit conifer seedlings if they
<br />exclude more competitive vegetation, such as shrubs
<br />(Amaranthue and others 1993, McDonald 1986).
<br />Once conifer seedlings are well established, grass
<br />cover is less detrimental to their growth than shrub
<br />competition (McDonald and Oliver 1984, McDonald
<br />1986). If grass cover is not too thick, it can potentially
<br />benefit tree seedlings. Green (1990) observed over 90
<br />percent survival of Douglaa fir seedlings on a site
<br />seeded after fire with cereal rye (Secale cereals). The
<br />rye, at a density of 9 plants ft_2 (100 plants m 2),
<br />provided shade to the seedlings during the first year
<br />after fire. The rye decreased to leas than 3 plants ft-2
<br />(33 plants m 2) the second year and essentially disap-
<br />peared in the third.
<br />Cereal grains such as barley, cereal rye, oats (Avena
<br />sativa), and winter wheat {Tritium aestiwm) appear
<br />to show great promise for producing cover that does
<br />not persist. Annual ryegrass was expected to behave
<br />this way, but it proved persistent beyond a wuple of
<br />years in some ecosystems (Barro and Conard 1987,
<br />Griffith 1998) and often produces maximum cover the
<br />second year after fire, ratherthan the first (Beyera and
<br />others 1998; compare Janicki 1989 with Conard and
<br />others 1991). A few reports cited initial concerns over
<br />the impacts of cereal grains on native regeneration
<br />that disappeared after further monitoring (e.g.,
<br />Callahan and Baker 1997, Hanes and Callahan 1995,
<br />1996, VanZuuk 1997). Somecereal grains mayexhibit
<br />allelopathy, inlubitingcompetingplant growth chemi-
<br />cally (Went and others 1952), but this has not been
<br />investigated under Seld conditions. Clearly more re-
<br />search on and monitoring of poatfire cereal grain
<br />seeding is needed, especially regarding the impacts on
<br />native herbaceous plants and conifer seedlings.
<br />In many cases natural regeneration provided as
<br />much cover as seeded species during the first years
<br />after fire, but good methods for assessing native seed
<br />bankviability are lacking (Isle 1998, Loftin and others
<br />1998). One standard teat for seed bank viability only
<br />identifies large-seeded species (by sieving them from
<br />poatfire soil samples) or those that germinate quickly
<br />(7 to 10 daygreenhouae germination test) (Dyer 1995).
<br />Species that will proride cover later in the winter or in
<br />the aecondgrowingaeason-the same time that seeded
<br />grasses provide most of their cover-are not detected
<br />by this method if they have tiny seeds or cold require-
<br />ments for germination. Better understanding of the
<br />natural range of vegetation response to fire would
<br />increase our ability to predict whether seeding is
<br />really necessary (Loftin and others 1998, Tyrre11981).
<br />Several interviewees suggested that more flexibil-
<br />ityinchooeing eeedmixeabeallowed inBAERprojecta,
<br />including the use of quick-growing annuals for ero-
<br />aioncontrol and slower growing native perennials For
<br />long-term ecosystem restoration, particularly native
<br />range. At present, BAER guidelines stress the use of
<br />only proven erosion-control species for emergency
<br />USDA Forest Service Gen. Tech. Rep. RMRS-GTR~3. 2000 51
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