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annuals, ouch as cereal grains or sterile hybrids, that <br />provide quick cover and then die out to let native <br />vegetation reoccupy the site. <br />Chaparrai.• Chaparral is the shrub-dominated veg- <br />etation type abundant in the low to middle elevation <br />foothills in California and the Southwestern States <br />(Cooper 1922, Keeley and Keeley 1988). Chaparral <br />stands are often located on steep slopes, burn with <br />generally high intensity, and typically develop water- <br />repellent soils. They become candidates for poatfire <br />seeding due to the threat of increased runoff and <br />sediment movement (Ruby 1989). <br />Concern over impacts ofpostfire seedinghas focused <br />on chaparral ecosystems because a specialized annual <br />flora takes advantage of the light, space, and soil <br />nutrients available after fire (Keeley and others 1981, <br />Sweeney 1956). Some of the dominant shrub species <br />regenerate after fire only from seed (Keeley 1991, <br />Sampson 1944). Moat published research on chaparral <br />comes from California (tables 6 and 7). <br />Bruahfields prone to fire and erosion occur at the <br />urban/wildland interface, where growing population <br />centers in lowland valleys have encroached on foot- <br />hillsand steep mountain fronts. The societal impacts <br />of wildfire and subsequent accelerated erosion in <br />California chaparral are enormous, as are the prea- <br />aurea to treat burned hillsides with grass seed to <br />protect life and property (Arndt 1979, Gibbons 1995). <br />Foresters in southern California began seeding <br />burned-over slopes with native shrubs in the 1920'x. <br />After finding that seeded shrubs emerged no earlier <br />than natural regeneration (Department of Forester <br />and Fire Warden 1985), they experimented with intro- <br />duced herbaceous epeciea such as Mediterranean <br />mustards in the 1930's and 1940's (Gleason 1947). <br />Mustards proved to be unpopular weeds with downalope <br />orchardists and suburbanites, so other species were <br />tested, including native and non-native eubshruba <br />and non-native grasses (Department of Forester and <br />Fire Warden 1985). By the late 1940'a annual ryegrase <br />(Latium muitifiorum, also called Italian ryegrasa), a <br />native of temperate Europe and Asia, had became the <br />primary epeciea used For poatfire seeding. Like mus- <br />tard, it was inexpensive, could be broadcast easily <br />from aircraft, was available in large quantities, and its <br />fibrous mot system appeared effective at stabilizing <br />surface soil (Barro and Conaad 1987). <br />The effectiveness of broadcast grass seeding for <br />erosion control on steep chaparral slopes has been <br />questioned (Conrad 1979), but relatively few data an <br />erosion response exist. The first watershed-scale reha- <br />bilitation experiment was set up at the San Dimas <br />Experimental Forest after a wildfire in 1960, includ- <br />ing annual and perennial grass seeding. The fret <br />winter after the fire was one of the driest on record <br />with negligible grass establishment (Corbett and Green <br />1965). The treatments were reseeded, and the next <br />year seeded grasses did not affect peak etreamflow <br />during four recorded storm events. The high-rate <br />annual grass treatment produced 8 percent grass <br />cover by the time of the last large storm event and <br />resulted in a 16 pement reduction in sediment produc- <br />tion over the season (Krammes and Hill 1963). Con- <br />tour planting of barley, which included hand-hoed <br />rows and fertilization, had the greatest impact on <br />sediment production (Rice and others 1965). Atl seeded <br />treatments had lower cover of native plants than <br />unneeded controls (Corbett and Green 1965). <br />Data collected by the California Department of <br />Forestry showed that ryegrass establishment was <br />typically poor in interioraouthern California and more <br />successful in cooler, northern or coastal locations <br />(Blanford and Gunter 1972). An inverse relationship <br />between ryegraes cover and native herbaceous plant <br />cover was observed, and Blanford and Gunter (1972) <br />felt that more data were needed to properly evaluate <br />the competitive effects of seeded ryegrasa on native <br />herbs. Range improvement studies found that high <br />seeded grass cover could reduce shrub seedling den- <br />sity (Schulz and others 1955). Blanford and Gunter <br />(1972) did not observe major failure of shrub regenera- <br />tion,though noquantitative measurementsweremade. <br />Ageneralnegativerelationehipbetweenryegrasscover <br />and erosion was observed using erosion pine. Blanford <br />and Gunter (1972), like Krammes and Hill (1963) and <br />Rice and others (1965), concluded that poatfire annual <br />grass seeding was an appropriate rehabilitation <br />method because its low cost made occasional seeding <br />failure an acceptable risk. <br />Cover or biomass of native chaparral vegetation, <br />especially herbaceous species, tended to be lower on <br />plots with high ryegrasa cover, both in operationally <br />seeded areas (Keeley and others 1981, Nadkarni and <br />Odion 1986) and on hand-seeded experimental plots <br />(Gautier 1983, Taskey and others 1989). Native plant <br />species richness was lower on plots centaining ryegrasa <br />(Nadkarni and Odion 1956, Taskey and others 1989). <br />Gautier (1983) and Taskey and others (1989) found <br />lower density of shrub seedlings, especially epeciea <br />killed by fire, on needed plots, and warned that long- <br />term chaparral epeciea composition could potentially <br />be affected by grass seeding. Taskey and others (1989) <br />also noted bare areas appearing in seeded plots where <br />ryegrasa died out after 3 years, resultingin lower cover <br />than on unseeded plots. These studies suggested that <br />ryegrase grows at the expense of native vegetation. <br />During a year in which total rainfall was exception- <br />alIy high compared to average, Gautier (1983) mea- <br />sured lees erosion from plots in which ryegraea seed- <br />ing increased total plant cover. On the other hand, <br />Taskey and others (1989) found no effect of ryegrase <br />on first-year postfire erosion with average rainfall and <br />12 USDA Foresl Service Gan. Tech. Rep. RMRS-GTR-63.2000 <br />