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i• <br />maturation. If this is true. then access to these sheltered off-channel pools may be ver}~ important <br />to successtul spawning in the upper reaches of the Colorado Ricer. Historically, bottomlands <br />that routinely flooded during the spring runoff period would have provided these warm <br />productive habitats; in recent years, flooded gravel pits may have provided the only compazable <br />habitat. <br />Razorback Sucker <br />The razorback sucker. an endemic species unique to the Colorado River Basin, w•as historically <br />abundant and widely distributed w5thin warmwater reaches throughout the Colorado River Basin. <br />Historically, razorback suckers were found in the main stem Colorado River and major <br />tributaries in Arizona, California, Colorado, Nevada, New Mexico, Utah, Wyoming, and in <br />Mexico (Ellis 1914; Hinckley 1983). Bestgen (1990) reported that this species was once so <br />numerous that it was cotrmtonly used as food by early settlers and, further, that commercially <br />mazketable quantities were caught in Arizona as recently as 1949. In the Upper Basin, razorback <br />suckers were reported in the Green River to be very abundant neaz Green River, Utah, in the late <br />1800's (Jordan 1891). An account in Osmundson and Kaeding (1989) reported that residents <br />living along the Colorado River near Clifton, Colorado, observed several thousand razorback <br />• suckers during spring runoff in the 1930's and early 1940's. In the San Juan River drainage, <br />Platania and Young (1989) relayed historical accounts of razorback. suckers ascending the <br />Animas River to Durango, Colorado, azound the tum of the century. <br />A mazked decline in populations of razorback suckers can be attributed to construction of damp <br />and reservoirs, introduction of nonnative fishes, and removal of lazge quantities of water from the <br />Colorado River system. Dams on the main stem Colorado River and its major tributaries have <br />segmented the river system and drastically altered flows, temperatures, and channel <br />geomorphology. Major changes in species composition have occurred due to the introduction of <br />numerous nonnative fishes, many of which have thrived due to man-induced changes to the <br />natural riverine system. <br />The current distribution and abundance of the razorback sucker have been significantly reduced <br />throughout the Colorado River system (McAda 1987; McAda and Wydoski 1980; Holden and <br />Stalnaker 1975; Hinckley 1983; Ivfazsh and Hinckley 1989; Tyus 1987). The only substantial <br />population of razorback suckers remaining, made up entirely of old adults (McCarthy and <br />Hinckley 1987), is found in Lake Mohave; however, they do not appeaz to be successfully <br />recruiting. While limited numbers of razorback suckers persist in other locations in the Lower <br />Colorado River, they aze considered rare or incidental and may be continuing to decline. <br />In the Upper Basin, above Glen Can}•on Dam, razorback suckers are found in limited numbers in <br />both len:ic and lotic eavironn:ents. Tile ;~: gest population o: razorback suckers ir. the upper <br />Basin is found in the upper Green River and lower Yampa River (Tyus 1987). Lanigan and Tyus <br />(1939) estimated that from 758 to 1,138 razorback suckers inhabit the upper Green River. In the <br />Colorado River, most razorback suckers occur in the Grand Valley azea near Grand Junction, <br />