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6 • <br />of its characteristics in a laboraton• flume stud}'. O'Brien (198-1) concluded that incipient motion <br />of the cobble bed is required to clean cobbles for spawning and estimated that this takes <br />discharges of about 21,00 cfs. However, Harvey et al.(1993) concluded that since flows <br />required for incipient motion of bed material aze raze (20 }•ear return period event) and spawning <br />occurs annually, another process must be cleaning the cobbles. Their study found that in Yampa <br />Canyon recessional flows routinely dissect gravel bazs and thereby produce tertiary bazs of clean <br />cobble at the base of the riffles. These tertiary bars are used by Colorado pikeminnow for <br />spawning. Harvey et al. (1993) recognizes the importance of high magnitude, low frequency <br />dischazges for the formation and maintenance of midchannel bars. Dissection of bazs without <br />redeposition by high magnitude flows would lead to conditions where spawning habitat is no <br />longer available (Harvey et al. 1993). <br />It is unknown whether tertiary bazs used for Colorado pikeminnow spawning in Yampa Canyon <br />aze available in the 15-mile reach of the Colorado River. There, significant motion of bed <br />material occurs at neaz bankfull discharge of 22,000 cfs (Van Steeter 1996). These flows occur <br />on average once in 4 years. Van Steeter (1996) concludes that flows of this magnitude are <br />important because they generally remove fine sediment from the gravel matrix which maintains <br />the invertebrate community and cleans spawning substrate. <br />Although the location of spawning azeas in the Colorado River is not well defined, the presence • <br />of larvae downstream of the Walker Wildlife Area, in the Loma to Black Rocks reach and neaz <br />the confluence of the Dolores River, demonstrates that spawning does occur. Osmundson and . <br />Kaeding (1989 and 1991) reported that water temperatures in the Colorado River were suitable <br />for spawning in the Grand Junction area. In 1986, a yeaz of high runoff, suitable temperatures for <br />spawning (20° C) occurred in the first week of August. In 1989, a year of low runoff, the mean <br />temperature reached 20° C during the last week of June. Miller et al. (1982) and Archer et al. <br />(1986) demonstrated that Colorado pikeminnow often migrate considerable distances to spawn in <br />the Green and Yampa Rivers, and similaz movement has been noted in_the main stem Colorado <br />River. <br />Miller et al. (1982) concluded from collections of larvae and young-of--year below known <br />spawning sites that there is a downstream drift of larval Colorado pikeminnow following <br />hatching: Extensive studies in the Yampa and upper Green Rivers have demonstrated <br />downstream distribution of young Colorado pikeminnow from known spawning areas (Archer et <br />al. 1986; Haynes et al. 1985). Miller et al. (1982) also found that young-of--year Colorado <br />pikeminnow, from late summer through fall, preferred natural backwater areas of zero velocity <br />and less than 1.5-foot depth over a silt substrate. Juvenile Colorado pikeminnow habitat <br />preferences aze similar to that ofyoung-of--year fish, but they appear to be mobile and more <br />tolerant of lotic conditions aw~av from the sheltered backwater environment. <br />Information onradio-tagged adult Colorado pikeminno~v during fall suggests that fish seek out • <br />deepwater azeas in the Colorado River (Miller et al. 1982), as do man}• other riverine species. <br />