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<br />.. <br /> <br />... <br /> <br />S+anForo + lt~an:J <br />I q~(pb- <br /> <br />.A.~, <br /> <br />__0 t"/r")~6 r <br />0-1- CL41 +-tJ vd <br />qL WcJ Y"4 <br />)q9~h- <br /> <br />9B. Fish of the Colorado system <br /> <br />1. A. Stanford &1. V. Ward <br /> <br />Introduction <br /> <br />The abundant remains of large minnows (Cyprinidae) in caves near desert <br />segments of the Colorado system (Plate Ic, e; cf. Minckley 1973), suggest that <br />the riverine fish were important sustenance for prehistoric man, The river <br />continues to supply sustenance, but the modes of transfer have changed dra- <br />matically. The vicarious regulation of the Colorado River, coupled with many <br />introductions of non-native species, has caused precipitous declines among the <br />indigenous fish, <br />This chapter describes the indigenous fish fauna and its relation to the <br />physiographic and hydrographic history of the basin (Stanford & Ward 1986a: <br />Fig. 1), and includes consideration of species in parts of the Colorado system <br />affected by desertification (e.g. Pluvial White River, Railroad Valley: Hubbs & <br />Miller 1948). We also indicate the status of the native fish in the regulated river, <br />and discuss the impacts of exotic species. <br /> <br />The native fish fauna <br /> <br />Thirty-two species of fish in seven families are indigenous to the Colorado <br />system (Table 1). Several have diversified in isolated local or sub-basin popu- <br />lations. Thus, four forms of the Colorado River chub (Gila robusta) are recog- <br />nised in the Gila, Virgin and Pluvial White rivers (Rinne 1976; Smith et al. <br />1979). Two subspecies of the spinedace (Lepidomeda mollispinis) are isolated in <br />the Virgin River and Meadow Valley Wash (Lee et al. 1980). Minckley (1973) <br />listed four subspecies of the desert sucker (Catostomus clarki) in Lower Basin <br />tributaries. Five subspecies of White River springfish (Crenichthys baileyi) are <br />isolated in desert springs in the Pluvial White and Moapa drainages (Deacon & <br />Bradley 1976; Williams & Wilde 1981), The speckled dace (Rhinichthys osculus) <br />is the most widespread native species, but populations in different sub-basins <br /> <br />The Ecology of River Systems, edited by 8, R, Davies & K. F. Walker <br />i] /986. Dr W. Junk Publishers, Dordrecht, The Netherlands <br /> <br />~ <br /> <br />.- <br />~ <br />t <br />--< <br />~ <br />~~,.... <br />, .. ..... <br />~ 0 ~, <br />'V~, <br />.... 7 :. <br />.;..~ <br />~ <br /> <br />385 <br />