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<br />002422 <br /> <br />Figure 2, Political and Ecological Boarders <br />in the United States and Canada <br /> <br /> <br />~~ <br /> <br />. <br /> <br /> <br />Political Map <br /> <br />tional approach. BIOGEOGRAPHY 57 (C.V. Mosby <br />& Co. 1983). E. Mayr declared that "the range <br />of a species is delimited by a line beyond which <br />the selective factors of the environment prevem <br />successful reproduction." POPULATIONS, SPECIES, <br />AND EVOUITIoN 30 I (Harvard Univ. Press 1970). <br />This would generate a boundary somewhat <br />higher than that of Brown and Gibson. M.D.F. <br />Udvardy had earlier restricted the boundary even <br />more by assening that "the limit of a well.doc- <br />umented area should connect those extreme <br />breeding stations which are not fanher from one <br />another than the distance that can regularly be <br />covered by normally dispersing individuals of <br />the species." DYNAMIC ZOOGEOGRAPHY 164 (Van <br />Nostrand Reinhold 1969). R. H. MacAnhur noted <br />that "we could also . . . define the nonhero <br />edge of the range as the point where the pop. <br />ulation density falls to, say, one.twentieth of its <br />density in the middle of the range or one-twen- <br />tieth of its maximum density." GEOGRAPHiCAl <br />ECOLOGY 146 (Harper & Row 1972). This def- <br />inition might draw a line well up Figure 1. <br />At present we (end to use the systematic <br />biology or, at most, the Brown and Gibson def. <br />inition of range. The practical problem with the <br />use of these more liberal definitions is that they <br />include patchy fringe populations within the <br />range boundaries. This means that the inference <br />of human. caused breaks in a species' distribu. <br />tion near its range limits can be drawn only from <br />a comparison of the existing situation (Q the <br />natural pattern of patchiness, and data to de. <br />scribe that natural pattern seldom exist. The only <br />way to avoid this problem is to define the range <br /> <br />. <br /> <br /> <br />Biological Map <br /> <br />boundary more conservatively, and that neces- <br />sarily means that some fringe locations capable <br />of suppotting a species will be excluded. <br />Resolution of this problem wi 11 not be triv- <br />ial. It must begin with political insistence that <br />the scientific community address the question <br />of defining a species' range. It should conclude <br />at leas! with a thoughtful set of rules by FWS. <br /> <br />Political Boundaries versus <br /> <br />Species Boundaries <br /> <br />The gray wolf is one of the more contro- <br />versial entries on the list of threatened and en- <br />dangered species. Where it exists in Minnesota <br />and Wisconsin it has caused endless bickering. <br />Where it is penetrating the northern Rockies, it <br />has precipitated a major political battle. All of <br />this has occurred despite the fact that if you ask <br />nearly any group of professional wildlife biol- <br />ogists if the wolf is threatened with extinction, <br />their answer will be a resounding no! Wolves <br />currently number fifty thousand to sixty thou- <br />sand animals and occupy the vast majority of <br />their original range in Canada and Alaska. The <br />clear fact, acknowledged by all sides in this dis- <br />pute, is that the fight over the wolf is a fight <br />over symbols or land use; it is not a fight to <br />preserve a species at risk of extinction. <br />To understand why we have embarked on <br />such a policy, compare a political map of the <br />United States and Canada with an ecological map <br />of the major biomes (Figure 2). The processes <br />Continued on page 58 <br /> <br />NR&E/SUMMER 1993 <br /> <br />The fight over the <br />gray wolf is a fight <br />over symbols or <br />land use, not a fight <br />to preserve a species <br />at risk of <br />extinction. <br /> <br />9 <br />