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<br />mate!y the same ttme every year, significant prob- <br />lems could arise if the high stress period was <br />extended late enough into the summer period by <br />increased snowpack. These phenologic events current- <br />ly oocur after the period of high water stress. If <br />they occurred during the period of high stress. a <br />significant growth reduction might occur. Further <br />coordination with the Tree Biomass and Tree Phenology <br />projects will be necessary to examine these implica- <br />tions. <br /> <br />4.5.4. Proposed Future Research <br /> <br />Little additional field data will be collected during <br />the upcoming year except to fill some gaps of missi~ <br />information. The majority of effort.will be concen- <br />trated on computer analysis of data for all six <br />plots and analysis and interpretation of the root <br />resistance study. When the tree core information <br />becomes available, it will be correlated with <br />moisture stress. <br /> <br />4.5.5 SummarY <br /> <br />The growlng,season of Engelmann spruce can be divided <br />into two parts concerning water s.ress, the snowmelt <br />period and the midsummer peribd. Water stress <br />during the midsummer period is primarily determined <br />by daily weather and expressed as two factors, one <br />atmospheric and the other deep soil. A regression <br />model was developed to predict water stress during <br />this period from the following variables (R2 - .81); <br />surface soil temperature, air temperature, -25 soil <br />temperature. and -15. cm soil water. <br /> <br />The snowmelt period is characterized by high levels <br />of ~idday water stress which become less severe as <br />snowmelt prQceeds and more soil is bared -of snow. <br />The 50% reduction in midday stress over this period <br />1s probably due to lowered root resistance to water <br />flow c8used'by increasing soil temperatures. This <br />theory is substantiated by preliminary analysis of <br />root resistance studies which demonstrated a rate of <br />water movement through roots at 7.5oC about 3X <br />greater than at O. SoC. ~ <br /> <br />The major effect of these high stress levels during <br />snowmelt would be in photosynthesis reduction rather <br />than cell division since trees were able to rehydrate <br />at night. ~s long as this high stress period occurs <br />before the beginning of major phenological activity, <br />little change in growth would be expected. If <br />delayed snowmelt from snowpack augmentations resulted <br />in the high stress period being extended later into <br />the year, occurrence of phenological events during <br />this stress period could result in a significant <br />alteration ~f growth pattern. <br /> <br />4.5.6. Literature Cited <br /> <br />Evans, A.K. 1973. Patterns of water stress in <br />Engelmann spruce. M.S. Thesis. Colorado State <br />University. 57 pp. <br /> <br />A cold tempera- <br />technique. <br /> <br />Evans, A.K. and C.P.P. Reid. 1974. <br />ture error in the pressure chamber <br />Can. J. Forest Res. (In press). <br /> <br />Reid, C.P.P. and A.K. Evans. 1971. Moisture stress <br />as an indicator of the effects of snow on biomass <br />product~on of forest trees, pp. 58-78. ~ An <br />evaluat~on of the ecological impact of weather <br />modification in the Upper Colorado River Basin. <br />Teller, H.L., J.D. Ives, and H.W. Steinhoff (eds). <br />CSU-DWS :Report No. 7052. <br /> <br />Reid, C.P.P. and Kent Evans. 1973. Moisture stress <br />as an indicator of the ef ects of snow on biomass <br />,production of forest tree pp. 34-39. In <br />Teller, H.L., J.D. lves, nd H.W. Steinhoff (eds). <br />CSU-DWS Report No. 7052-Z <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I; <br /> <br />27 <br /> <br />I~ <br />i <br />, <br />Ij <br /> <br />I <br />11 <br />1 <br />Ij <br />II <br />j <br />II <br /> <br />j <br />