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<br />OJn742 <br /> <br />" <br /> <br />that biogeographical features (latitude, altitude, slope, <br />distance from Pacific Ocean) and hydrological vari- <br />ables (current velocity, mean depth) were most useful <br />in delineating site groupings in a survey of streams <br />in north-western North America, Within basins, <br />Ormerod & Edwards (1987) determined that water <br />chemistry (pH or total hardness) and associated geo- <br />logy were the dominant correlates with macroinvert- <br />ebrate assemblages, along with slope or distance from <br />source in the River Wye, Richards et ai, (1993) reported <br />that substrate variables were the most important fac- <br />tors explaining variation in macroinvertebrate commu- <br />nities in a Michigan agricultural watershed, As <br />demonstrated by all these studies, the type of environ- <br />mental factors affecting invertebrate community distri- <br />bution in stream ecosystems differs based on the <br />spatial scale (regional, basin, within stream) examined <br />and the large-scale environmental gradients (e,g, alti- <br />tude) present within the study area, For the South <br />Platte River Basin, ecoregions and physical (geomor- <br />phology) and chemical factors (as affected by land <br />use) within ecoregions are dominant factors affecting <br />invertebrate distribution. <br /> <br />Acknowledgments <br /> <br />" <br /> <br />This study was supported by the US Geological <br />Survey's National Water Quality Assessment Program: <br />South Platte River Study, We thank KF. Dennehy, J,R <br />Ortiz, RC. Averett, V. Averett, RC. Petersen, LR <br />DeWeese, P,B. McMahon, D.W, Litke, J,E, Paschal, RB. <br />Swanson, GD, McElhany, and KL Findell for field <br />assistance, We thank H,V. Leland for assistance in data <br />analyses, We thank H,V. Leland, KF. Dennehy, R,G, <br />Smegl, T,M, Short and an anonymous referee for <br />their critical review of the manuscript. This paper is <br />dedicated to the memory of Robert C. Averett <br />(deceased June 25, 1993) whose leadership in aquatic <br />ecology and inspiration to aquatic ecologists will <br />be missed, <br /> <br />References <br /> <br />Anderson JR, Hardy E,E" Roach J,T, & Witmer RE, <br />(1976) A land use and land cover classification system <br />for use with remote sensor data. U.S. Geological Survey <br />Professional Paper 963, 28 p, <br />Benke AC., Van Arsdall T,C., Gillespie D,M, & Parrish <br />EK. 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