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<br />~ <br />~ <br />Q <br />~ <br />o <br />~ <br />~ <br />~ <br />.. <br />~ <br /> <br /> <br />, <br /> <br />SI'RL\KBA..'nt <br /> <br />, - <br /> <br />. <br /> <br />7 <br />. <br />,8 <br />11 <br />12 <br /> <br />'00 <br />CHA.'rnEL WIDTH <br /> <br />. p . Prl~ry Production; R . Ecosystem Respiration <br /> <br />j'9 <br /> <br />- <br /> Organic FIR- Heterotrophic <br /> Matter ~AT" ., <br /> Input. Autotrophic <br />{Allochthono\lll) <br /> INPt.'TS <br />C1'OM FPOM <br />IUgh 1.0>. <br /> <1 Hetecrot'fophlc <br />Lov MiUh <br />(Reduc- <br />.d) <br /> >1 Autotrophic <br />-- <br />Low H:lgh <br /> <1 I lie tertrophlc <br /> I -- <br /> <br />Prl~ry <br />Production <br /> <br />INPti'S <br /> <br />Low <br /> <br />(Algae aod <br />Mosus) <br /> <br />Periphyton <br /> <br />Bigh <br /> <br />{Algae and <br />Hydroph:nes} <br /> <br />Perlphyton <br /> <br />~ Low <br />r-" <br />, _ (Primarilv <br />Planktonic) <br /> <br />Figure 3. Comparhon of the biological habitat of small, medlU1:1 and large sereaa aCCI~rdln& to <br />the River Continuum Concept. <br /> <br />being cooler in the summer months, warmer in the wint.er months and having <br />lower seasonal and diurnal temperature variation. <br /> <br />Streams with closed riparian canopies characteristically receive <br />significant amounts of coarse particulate organic mat.ter (CPOM) in the form <br />of leaves. needles, flowers, twigs, and branches. In many st reams, the <br />major influx of CPOM occurs during the annual autumn leaf fall. The ratio <br />of gross primary production (p) to community respiration (R) is normally <br />less than one over most of the annual cycle indicating that headwater <br />ecosystems are strongly heterotrophic. ThUll t.he maj or portion of the <br />trophic resource is imported from the adjac.mt ecosystem in the form of <br />CPOM during the fall. <br /> <br />Medium-Size Streams <br /> <br />Medium-size streams in the range of fourth to sixth order are <br />distinguished from small headwater streams and large streams by a number of <br />easily observable characteristics (Figure 2)" As small tributaries uni te, <br />channels progressively increase in size becoming wider and deeper. The <br />gradient or slope is generally less than headwater reaches. Volum" of <br />water flow increases significantly with the Increased tributary input. The <br />dominant substrate type is related to watershed geology, but <br />characteristically ranges from coarse to fine particles represented by <br />