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<br />62 <br /> <br />saltgrass (Distichlis sp.), seepwillolJ (Baccharis sp,.), and sycamore <br />(Platanus sp.). The most important water--consuming dparian phr<!atophytes <br />in the western United States are cottonwood, willow ,md salt cedar. <br />Robinson (1958) estimated that over 6.5 million ha (16 million aeres) of <br />phreatophytes in the western states annually transpire over 31,000 milllon <br />m3 (25 million acre-feet) of water. It was also estImated that the <br />aggressive saltcedar occupies over 526,000 ha (1.3 mj,ll1on acres) <br />transpiring over 6 million m3 (5 million acre-feet) of water. WlLter use by <br />salt cedar was found to vary from 0.61 m to 2.4 m (2 to 8 feet) pe,r year, <br />depending on location, density, salinity, and depth of the water table. <br /> <br />Riparian Ecosystem Succession <br /> <br />The riparian zone and its fluvial system are dynamic and <br />interdependent. As a stream overflows its banks, natural levees form from <br />the deposited sediments. Overbank flows supply water to adjacent <br />floodplain wetlands which store excess runoff water. Additionally, <br />overbank flows may facilitate groundwater recharge. As a stream erodes its <br />channel on the convex curve of a bend and deposits sediment on the concave <br />side, the meander moves laterally and down the valley while maintaining <br />constant overall channel shape and size. These two processes, overbank <br />flow/deposition and lateral migration, are the most significant forces in <br />formation of a floodplain. The riparian zone responds by establishing <br />successional stages (sere) of plant growth on new sand and gravel bars, <br />often beginning with sedges, rushes, willow (i.e. Salix spp.) and other low <br />plant cover. As gravel bar communities become more distant from the <br />shifting channel, they begin to stabilize ,a,nd are further colonized by tree <br />saplings. Gradually, a new riparian zone is established. <br /> <br />,.,:J;~eprinciples and terminology of ecological "su,ccession" apply to <br />riparian ecosystems and thereby facilitate understandLng, description and <br />prediction of outcomes from catastrophic events. Rip,arian ecosystem <br />"succession" is an orderly process of community development that <br />(1) involves changes in species stt'Ucture lind commun1:ty processes with <br />tille, (2) results from IIIOdif1catioll of the phySical el2vironment b:r the <br />C_nitY'and (3) cullll1nates in' a jltabilizl!d ecosy.tell in which maximum <br />biomass and symbiotic function bet..een orgunislDS are maintained P,!r unit of <br />available energy flow (Odum 1971). The sequence of communities w:lthin the <br />riparian zone is called the "sere, - and thE' relati vel)' transi tory <br />communities are called "seral stages." <br /> <br />A final, stable, riparian co~nity in a developoaental serie!l (sere) <br />is called the "cliillax." In the ab6ence of perturbation, the comllltlnity is <br />self-perpetuating and in equilibrillm with the physical environment:. Odum' <br />(1971) suggested that two types of climaxes be recognhed; II ".E!!..utic <br />clillax" which is in equilibriulI wiC;h the I.ural cl1lDlllte and towards which <br />all .ucceastonal development in a' given reach is tend1,ng and the "edapt1c <br />cl1eu:- Which is modified by local conditions of the s:ubstrate. Succession <br />may end in an edaptic climax where hydrological conditions, climate, soil, <br />topography, floods, fire, drought or other disturbances are such that the <br />climatic climax cannot develop. <br />