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<br />56 <br /> <br />~iparian Ecosystems and the Edge Effect <br /> <br />The large nWllber of vertical and horizontal strlltll caused by riparian <br />zone vegetation and the associated high productivity, l;pecies diversity and <br />densitj"are collectively known as the "edge effect" (FIgure 2). Those <br />species which occur primarily, are 1Il0st-aiiUndant or spE!lld the greatest <br />amount of time in the ecotone are called "ed.~ species." <br /> <br />r.;,!ii~. <br />\/(CnllTlVi!Sll'ljl"" ~~~ <br /> <br />l!~f ''''-?_ <br /> <br />2 F~-~~i;'" '", "" ~ ';'j'~ <br />1 [3/').,- -- "I'. t - - <br />, , , <br />~, ',~ " <br /> <br /> <br />1 <br /> <br />2 3 4 <br /> <br />5 <br /> <br />UK;f~ <br />f1..u,..1.n\pa""I'II_.trr..qu.""I'I'l~.""h;'l"""'''''''f'''''09''' <br />'l'Idll'..I.ln ',.C>IItP.....II...Ir.....H..'... ThiIP'Oc!\II:.'I'l.b,t..lI!W <br />. g,'.I., .......... 0' .ptel.., ...IIHII"9I1'I' ......,.1]>., pi"". 'II'- <br />c:",,,,",c__""',trvcl\lN <br /> <br />(Modified frolll ThOllla8 H d. 1979) <br /> <br />The dramatic contrasts between the vegetative complex of the riparian <br />zone and that of the adjoining aquatic and terrestrial zones add to the <br />structural diversity of the riparian co~~unity. Wetlands, ponds, oxbow <br />lakes, seeps, springs and minor drainages provide edges of stark contrast <br />to the surrounding semiarid or arid zones. Moreover, riparian zones <br />dominated by deciduous vegetation provide on,e, type of h,sbitat during <br />leaf-out aonths and another type of habitat dluring the 'dnter following <br />leaf-fall. The meandering shape and linear nature of the riparian zones <br />maximize the development of the edge effect and edge sp,ecies. Riparian <br />zones in semiarid and arid regions produce f,.,r more edg,"s within a small <br />area than would be expected or found in the ,adjacent sagebrush terrestrial <br />zones (Figure 3). <br /> <br />Functional Definition of Rip~!!an Vegetl~ <br /> <br />Riparian ecosystems are characterized bl' stream bank vegetation with <br />more hydric or mesic (i.e., moderately moist) growth habits than adjacent <br />, drier upland plant communities. All riparian plant comDlunities are <br />'--dependent upon high water tables or overbank flooding tel meet their <br />l10isture requirements. They may also depend on overbank flooding for the <br />deposition of substrates and nutrients necessary for regeneration, <br />establishment and maintenance. <br /> <br />Functionally, riparian ecosystemS strongly influence adjacent ..quatic <br />zones. To a large extent, the riparian zone regulates the productivity of <br />, the aquatic zone by supplying organic detritus in the form of leaves, <br />\~tems, twigs, branches, etc. to the stream. ~rhis has prompted SOme workers <br />