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<br />1:';\.LLoFQ,:!\1 i !lrAf.T~; ~1l ;:r,. :~.;rAtJ;.-,,!..L:!~'.1~:~I^'fI,t}.l~._""lli_1HE.2~.!'LJtAN MOUNTAINS OF COLORADO <br />C:HAJ:'TER 1 ~!i.!.RpI>UCJ ((IN <br /> <br />r" fI 1"':14, ,I, }~" SI:,"'IIf'.\', II, ....'. Stctnhnff. ,,"r1 W. W, Howell <br /> <br />Thill docUlnent tfi the flnnl l'I'pI'rt 111' tIL... Sail Juan <br />Ecology Project (S.Jf.P) \oIhJdl hl1s been supported by till' <br />UnitEld ~tRte6 BureAlI of ReeJ.lmlltiOIl unrter C('ntract <br />No. 14-06-1l-705:? (rom 1970 tn 197,. All 1~lIrl1er study. <br />that was prepRred n9 ;J prl1hlem l1nalysis, _has been re- <br />ported previnUJ'lly (S.lEI'. Phfls(' I FIIIIII It(")lnrt: TC'lJer <br />et R 1. 1970). 'I'll(' 111"NI~'1I1 r~pl1rt <PhI'tAf' II) llnl'ft nnt <br />include ony or th(' fOnr)y. I'r"pnrnl.lrv rnntr.rlnl. <br /> <br />SUMMARY OJ.' t:.F.Y f:(INCLl!S IONS <br /> <br />This Summar condenses the evaluat iOIl of ecologic <br />imD8ct8 due to v8rvin~ snoyPac in tie an uan oun-' <br />taina. It is based on a study of the influence of a <br />snowfall whlch Is quite vsriable in both space and <br />time. O1rect observation of the effects of snowpack <br />fugmentation were not possible during the 5-year life <br />of the San Juan Ecolo y Project. but the relationships <br />tlsta . e etween ecoaystem components an .now <br />conditions allow the eatinmtion of pOfll'llble cUr-cta of <br />long-term successful cloud Ap.cdlug. <br /> <br />The ststementA which fnllow trent thE' mOlln r.r.osyatem <br />components in urdt'r, Thr 111I1I(I(:t IIf ~llO""l'nrk HllH,ln('n- <br />tation on each ecosystem ,"umlt(lllenl IN nl'rnrlR('11 IIC~'lIrtl- <br />ing to the nLltun' tlf lllf' t"nvlr-onmcntnl ch^Il~p. which <br />produced the effect. The hAsic environmental changE'S <br />are the addition of more snow and more silver. Such <br />changes could 1nltjnt\~ n chain of effr.l:ts thnt would <br />radiate throughout tlw {'coaystem. The firat links in <br />the chain which havt! .'1 slR"ificant impact on ecosystem <br />components are: (I) Illwer Roil tempr.rAtllre In the <br />spring. (2) more moisture In the 8prin~. (3) deeper <br />snowpack. and (I.) more ~Jlver" <br /> <br />This summary includes only statements verifiable from <br />the data. with causal connections and conclusions <br />that would be acceDtcd by scientific peers in envlron- <br />, ment41 disciplines. Speculntiun about causal links in <br />the chain, or extension of impliCAtions beyond effects <br />which were studied in the chain, and where the extension <br />involves appU,cnt!on of Rene-rAl eC(llo(!,lc llndQr8J:..!!ndln~, <br />tire avoided in this summary. Such appropriate ec1en- <br />titie conjecture can be tound in Chapters II and lIT <br />and in seaments of individual reports that are spec- <br />ulative by implication t.hrollRh the use of terms such <br />88 "may be" or"pt"ubahly", nr whidl Art'; IlJbelled 1'1.8 <br />lib road sign1f1cnnce" Ill" "Implications." <br /> <br />Many other findings, which are not dlreC"tly related to <br />the effects of snoWf'ad augmentation on the, indicated <br />ecosystem components Bre a natural rart of such studies. <br />For a Bummary of theRe find1nR9 reference is made to <br />the abstracts of in(livJdIIU\ report~ Of to the detailR <br />of the individual fp.ports themse]vt's III ChApter IV. <br /> <br />~ <br /> <br />Onl twu (!nv:J rU,lllll'HI 111 ChilllWS ~'hl('h ml'ht he Jnduced <br />by increasel.l f1IHlW nil ha\'l.l Iwen !mllld Cf' r(.Jllte slgnif- <br />icantl to .Jantl:!.111 tilt" SOln ,1111111 N.llml1dnsi llllo'er <br />aoil temperatures lln nl(lrc so mu sture. <br /> <br />_ Lower SoiJ 'fempcrAtUTl'R <br /> <br />itlatlon of shoot cloll ,,,tinll .....,S d",'1.,yp.d for pl:mtA <br />oth in the tmlllrn ami fllrc9lH nl" 1\ rt>::IllJ t 'J l(l~'(.'r <br />"801 tem eral"llreS 8f1S01" nte w t 1;1 C!~pcr 9nOWl'lIC fur <br />',the species 8tudiC!d. Thefle il\.: II el .ng ernann spruc:" <br />Clli.!! .cnr.e!..m..!D.!l!-.!.J. (l'l<!,kil.t! 111''''':11 <l'E.!'.u_U'-s, ~'.I_tl?J!;I.~.~). <br />Thurber rttsr:ue (,,)..~_t:..UY.;1 1-'~\I.rE.('r',). 1\11.1 IlIlm.'rUlI!'I <br /> <br />19 <br /> <br />hcrhl1c:eoUR species in both the tundra and fOrf':fit meA- <br />dows. e dela vas most apparent in apecies~ith <br />Rrowth p,ointa n t e mate a e zone 0 ower- <br />tern eratures st the round level. This includes pri- <br />marily tie er aceous spec ell. e growth regions of <br />tref'!B, in the cambium and terminal bud8. WElre affp.ctl'i <br />morl' tndirectly, pn..sibly throuah 80il temperllturf! <br />e(fe-I:tA on root function. From this A les8 clt'Br re- <br />lntionehip between growth inception snd snowpack VB!:. <br />AnticJpAted although there "was a general correlation <br />uf bud buret dstes for both spruce and aspen with thl:! <br />snow clear dates. HQisture stre88 in spruce was un- <br />usually hiah durin a the period of snowmelt and decrees <br />dramatically at about the .now clear date. This did <br />not affect the annual total radial growth of spruce, <br />which initiated 2 to 4 weeks .earlier than bud burst, <br />apparently independent of snowpack. Resultant annual <br />biomass Rrowth of .pruee w...remarkablY constant ~ <br />year to year, unrelated to anowfall. Aspen was more <br />variRble in annual biomass product10n. Aspen growth <br />inrept ion wall dl!'layed by ,. later anow c!(OAr dllt~. bllt <br />we did not determine whether or not this waR directly <br />ltnked to variations in biomass production. <br /> <br />The delay in bednnltfR of growth persisted throuRh <br />thE' other stagea of the life cycle of mo~t plants, <br />'hut with less magnitude. There seemed universally a <br />compensatIon tactor Whlch rerm1~~ed every 8pecIes <br />to complete its annual eye . of development each <br />.summer. regsrd~e.. at anowpack depth. Except tor <br />spruce, there was no Slml~ar compensatory mechanism <br />for biomsRs production. In moat herbaceou9 species <br />of the tundra, biomass production was inversely <br />related to depth of snowpack. <br /> <br />No effect was found on either initiation of developmen <br />or biomass production of oak (Quercus Kambellii), pre- <br />sumAbly because even in a deep snow year the snow had <br />always disappeared from oak plata before the enrliest <br />date of growth initiation. <br /> <br />- tlnr.. Moisture <br /> <br />Ph)'tosociological 8tudies in both forests and tundra <br />showed that plant communities ware arranged along <br />8now clear date gradients. In the lore.ts ask give. <br />WR)' to aepen, then to .ubalplne fir (Ables lasiocarpa), <br />and finally to Englemann spruce a. the snow clear date <br />extends later into the year. The change was less <br />obvious in the drier communitles. t.e. ln oak. ,The <br />tundra changed from xeric (drier) to more mesic or <br />even hydric communities. This could be expected to <br />result in an increase in frequency of the rarer species <br /> <br />A dendrochronologie survey of pondeross pine (Pinus <br />ponderosa) immediately outside the target area-shOWed <br />it~ growth on dry sltes to be a.soctated with December <br />preripitation. The relationahip is a direct one and <br />Ine would reaurnabl ahow an increa.e in ro'Wth if <br />cloud seeding were to a to m -w nter snow 8 L 0 <br />'Buch relationship WIlS found for Engellllann spruce, the <br />(lull' other species on which dendrochronologlc work <br />has I>l!'!en conducted in thls study. <br /> <br />~nlf1l~" <br /> <br />Only the lo....er soil temperature and greater aDo\,,' depth. <br />which ~lRht be expected to follow an increase in snow- <br />fall. have been found influential on an1mal sctivity, <br /> <br />A-13-2 <br />