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Productivity (Number of Fledged Young and Fledging Rate) <br />Kirsch (1996) estimated fledglings per nest by defining the number of nesting pairs as the <br />maximum number of active nests and broods present at a colony during a single visit. <br />Kirsch (1996) recognizes that this assumes re- nesting pairs remained at their original <br />colony or that emigration and immigration of pairs must be equal. <br />NPPD, who provides data for the NGPC annual central Platte River census, expresses <br />fledge ratios as fledglings per nest. It is the empirical equation of all chicks documented <br />to fledge divided by the number of nests observed within a colony (Jenniges 2001). <br />Breeding Population Structure <br />Least Tern Breeding Population Structure <br />The least tern breeding population is comprised of metapopulations, which means that <br />breeding subpopulations occurring within local areas are indistinct components of the <br />larger breeding population (Wiens and Rothberry 1981, Pulliam 1988, Hanski and Gilpin <br />1991 in Kirsch and Sidle 1999). The bare sand areas used for nesting areas are <br />ephemeral, and the quality and amount in any one area can vary from year to year <br />depending on variable conditions that maintain habitat (Sidle et al. 1992). Consequently, <br />the tern breeding population and the reproduction rates on a particular portion of a river <br />system can fluctuate from year to year (Lingle 1993b) <br />Movements of individual breeding birds between nesting areas from year -to -year are <br />common. Least terns banded at central Platte River nesting colonies have been later <br />observed nesting up to 170 miles (273 km) from their banding origin (based on 163 <br />sightings of 109 individuals). Twenty -eight percent of the least terns returned to the <br />colony where they were banded as chicks. Movements have been documented between <br />the central Platte River nesting area and nesting areas at Quivira National Wildlife <br />Refuge (NWR), and the lower Platte River, near Fremont, Nebraska (Lingle 1993c in <br />FWS 1997). <br />Piping Plo ver Breeding Population Structure <br />Piping plovers are semi - colonial, and will nest in or near least tern colonies. Average <br />nesting colonies in Nebraska have 1 to 3 pairs of piping plovers (NGPC 1995). Piping <br />plovers frequently change territories following nest failure and between years. Piping <br />plovers banded at central Platte River nesting colonies have been later observed nesting <br />up to 125 miles (200km) from their banding origin (based on 71 sightings of 57 <br />individuals). Fourteen percent of re- sighted birds were banded as chicks. Thirty -two <br />percent of piping plovers returned to the colony where they were banded (Lingle 1993c <br />in FWS 1997). <br />Draft Baseline Report, July 2002 <br />4 -7 <br />