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2.4. Effect of Fires on Runoff <br />The effect of wild and prescribed fires on runoff is highly <br />variable because they can affect both the vegetation <br />canopy as well as the physical properties of the soil. At <br />one level, the effect of fires on runoff is similar to forest <br />harvest, as any reduction in the forest canopy will alter <br />the amount of interception and transpiration. Fires may <br />also consume some or all of the litter and duff layers, and <br />this also can increase runoff by reducing the total water <br />storage capacity at a given site (Kittredge, 1948). <br />The greater concern is the potential for high- severity <br />fires to alter the surface organic layer and mineral hori- <br />zons in ways that reduce infiltration and increase runoff <br />and erosion rates. The reduction in infiltration can oc- <br />cur by several processes. High - severity fires consume <br />all of the surface litter and duff. Sustained soil heating <br />can burn off much of the organic matter in the top few <br />centimeters of the mineral soil. In this situation there is <br />little protection of the soil surface from rainsplash, and <br />the disaggregated soil particles and ash can clog up the <br />larger soil pores that are crucial for maintaining the high <br />infiltration rates typical of forested areas (Terry and <br />Shakesby, 1993). <br />The second important change is the generation of a wa- <br />ter repellent (hydrophobic) layer at or slightly below the <br />soil surface. Fires can generate a water repellent layer <br />by volatilizing hydrophobic compounds in the organic <br />material, and some of these are driven downwards where <br />they condense on cooler soil particles at or below the <br />soil surface (DeBano, 1981; Letey, 2001). The amount <br />and type of these compounds varies with the type of <br />vegetation, while the depth at which these compounds <br />condense is a function of soil heating. In hotter, slower - <br />moving fires there is more soil heating and these volatile <br />compounds may be deposited at depths of 2 -6 inches be- <br />low the soil surface. In low- temperature and faster -mov- <br />ing fires there is less soil heating and these compounds <br />condense closer to the soil surface. Stronger water repel- <br />lent layers are associated with increasing burn severity, <br />as high- severity fires vaporize more organic compounds <br />and thereby generate a stronger and more continuous <br />water repellent layer (Tiedemann et al., 1979; DeBano, <br />1981). The development of a water repellent layer is of <br />concern because this can greatly reduce infiltration rates. <br />Once rainfall saturates the thin layer of ash or soil above <br />the water repellent layer, any additional rainfall will run <br />off as overland flow. <br />The development of a post -fire water repellent layer has <br />been most extensively studied in chaparral ecosystems, <br />17 <br />but post -fire water repellency has also been documented <br />under a number of different conifer species, including <br />ponderosa pine (Helvey, 1980), lodgepole pine (Meeu- <br />wig, 1971), and Douglas -fir (Helvey, 1980). Post -fire <br />water repellent layers are believed to be less likely in <br />vegetation types, such as aspen, that have less surface <br />fuels and fewer secondary compounds (MacDonald et <br />al., 2000). <br />Post -fire water repellency is generally believed to be <br />more severe in areas with coarse - textured soils. Coarse - <br />textured soils have a much lower particle surface area <br />than fine - textured soils, and this effectively results in a <br />greater concentration of the water repellent compounds <br />per unit surface area (Meeuwig, 1971; DeBano, 1981). <br />Changes in soil moisture will also affect the strength of <br />a water repellent layer. If moisture is present, a water <br />repellent soil will slowly wet up due to the strong hy- <br />draulic gradient and movement of water vapor (DeBano, <br />1981). As a water repellent soil wets up, there usually <br />is usually a threshold at which a soil ceases to be water <br />repellent (Crockford et al., 1991; Dekker and Ritsema, <br />1994; Doerr and Thomas, 2000). Upon drying the water <br />repellent conditions can be re- established ( Shakesby et <br />al., 1993). <br />The persistence of a post -fire hydrophobic layer will <br />depend on the initial strength and thickness of the hy- <br />drophobic layer and the animal activity, plant regrowth, <br />and physical and chemical processes that collectively act <br />to break down the hydrophobic layer (DeBano, 1981). <br />Soil water repellency usually returns to pre -burn condi- <br />tions in no more than six years (Dymess, 1976; DeBano, <br />1981), and several studies have documented a much <br />more rapid recovery (e.g., DeByle, 1973; Reeder and <br />Jurgensen, 1979). <br />Until recently there has been relatively little work on <br />the development and persistence of post -fire soil water <br />repellency in Colorado. In most cases the presence of a <br />water repellent layer has been inferred from the observed <br />post -fire increases in runoff and erosion (e.g., Moms and <br />Moses, 1987). However, in the summer of 2000 detailed <br />measurements of soil water repellency were made on <br />five fires in the Colorado Front Range that burned from 1 <br />to 22 months earlier (Huffman et al., 2001). Strong water <br />repellency was found in ponderosa and lodgepole pine <br />forests that burned at high or moderate severity, regard- <br />less of whether the fire was a wildfire or a prescribed <br />fire. Areas that burned at low severity generally had <br />little or no more water repellency than unburned areas. <br />Soil water repellency was strongest at the soil surface, <br />