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and freezing at the surface of the snowpack, and possibly <br />an increased persistence of the snowpack as a result of <br />the increased peak SWE. <br />Figure 2.2 showed the effect of forest harvest on the av- <br />erage annual hydrograph for the Fool Creek catchment <br />in the FEE On average, the annual daily maximum peak <br />flow increased by 23% as a result of removing 50% of <br />the forest cover (Troendle and King, 1985). The average <br />number of days with flow at or above bankfull increased <br />from 3.5 to 7 days per year (Troendle and Olsen, 1994). <br />A reanalysis of data from the Wagon Wheel Gap study <br />showed that the annual daily maximum flow increased <br />by 50% (Van Haveren, 1988), and a similar increase in <br />the size of the annual snowmelt peak occurred as a result <br />of the shelterwood cut on the North Fork of Deadhorse <br />Creek (Troendle and King, 1987). These and other stud- <br />ies indicate that complete removal of the forest canopy in <br />small watersheds in the sub - alpine zone will increase the <br />size of the annual daily maximum flow by about 40 %. If <br />only part of the forest canopy is removed, the increase <br />in the size of the annual daily maximum flow is directly <br />proportional to the amount of basal area removed. <br />Increases in the instantaneous annual maximum flow are <br />more variable (e.g., King, 1989). In general, harvest -in- <br />duced increases in the instantaneous annual maximum <br />flow should be similar to, or slightly smaller than, the <br />change in the annual daily maximum peak flow because <br />forest harvest will have a progressively smaller effect on <br />the highest peak flow (i.e., peak snowmelt is limited by <br />the amount of energy that can be delivered to the snow - <br />pack). For example, forest harvest had no detectable ef- <br />fect on the three largest instantaneous peak flows on Fool <br />Creek despite the significant increase in the annual daily <br />maximum peak flow (Troendle and Olsen, 1994). <br />Harvesting 24% of the 4,100 -acre Coon Creek water- <br />shed increased both the annual daily maximum and the <br />instantaneous peak flows resulted by about 8 %, but these <br />increases were not statistically significant (Troendle et <br />al., 2001). However, there was a significant increase in <br />the 701 to 901 flow quantiles, where the value nearest <br />zero represents the lowest flow on record and the value <br />closest to 100 represents the highest flow on record <br />(Troendle et al., 2001). The absence of a statistically <br />significant increase in the annual peak flows in the Coon <br />Creek study as compared to Fool Creek is partly due to <br />the much smaller number of pre- and post - treatment data <br />points, the variability in the relationship between the <br />treated and the control basins, and the fact that only 24% <br />of the Coon Creek basin was harvested. Peak flow in- <br />creases also may be smaller in larger basins with a wide <br />14 <br />range of elevations and aspects, as the peak flows from <br />different parts of the basin may not be synchronized. <br />The effect of forest harvest on the size of peak flows is <br />much more complicated and controversial in areas where <br />the largest peak flows are caused either by mid- winter <br />rain -on -snow (e.g., Harr, 1986) or rain-on- spring -snow- <br />melt (MacDonald and Hoffman, 1995). Since rain -on- <br />snow events are rarely encountered in Colorado and <br />don't appear to be a primary cause of the peak flows <br />above 7500 ft (Jarrett, 1993) or the largest runoff events <br />below 7500 ft (N. Doesken, Colorado State University, <br />pers. comm., 2000), the effect of forest management on <br />rain -on -snow events will not be considered further. <br />Some forest hydrologists have suggested that watershed - <br />scale increases in the size of peak flows can be exac- <br />erbated or ameliorated by harvesting different portions <br />of the basin to alter the timing of runoff from different <br />tributaries (e.g., Harr, 1981). The argument is that for- <br />est harvest close to the mouth of a basin may accelerate <br />runoff and help desynchronize the runoff from the lower <br />portion of a basin relative to the upper portion. Alterna- <br />tively, forest harvest in the upper portions might acceler- <br />ate the delivery of water to downstream areas and further <br />increase peak flows by synchronizing the peak flows <br />from the upper and lower portions of a basin. <br />Research in Colorado and other snowmelt- dominated <br />areas indicate that forest harvest usually has little or <br />no effect on the timing of the peak flows. Forest har- <br />vest caused no significant change in the timing of the <br />snowmelt peak at Wagon Wheel Gap, Deadhorse North, <br />Deadhorse South (Troendle and King, 1987; Troendle, <br />1987a), or Coon Creek (Troendle et al., 2001). In con- <br />trast, the annual maximum peak flow on Fool Creek <br />occurred an average of 7.5 days earlier (Troendle and <br />King, 1985). The earlier peak at Fool Creek is attributed <br />to the faster and earlier melt in the lower parts of the <br />catchment, and this superimposed an earlier and sharper <br />peak on a formerly flat or bimodal snowmelt peak (Fig- <br />ure 2.3). Overall, there is little evidence to suggest that <br />forest harvest can consistently and significantly affect <br />the timing of peak flows in Colorado. Hence the issue of <br />synchronization appears to be of little practical signifi- <br />cance, and will not be considered further. <br />2.2.6 Effects of vegetation change on low flows. <br />In contrast to changes in the size of peak flows, an in- <br />crease in the size of low flows is generally considered <br />beneficial, as the additional water may be useful for <br />water supply purposes, increasing instream habitat, or <br />improving water quality (MacDonald et al., 1991). Stud- <br />