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populations the chances of mating between relatives increase, and Fis may increase <br />accordingly (Frankham et al. 2002). The parameter FST is a measure of genetic distance <br />among populations. When populations are isolated (i.e. due to geographic barriers or <br />temporal differences), individuals within those populations mate with one another rather <br />than with individuals from other populations. The resulting population subdivision is a <br />form of non-random mating, and the FST value reflects how different the populations <br />have become as a result (Gillespie 1998). The parameter FIT can be thought of as the <br />overall measure of inbreeding. It is "the most inclusive measure of inbreeding in that it <br />takes into account both the effects of nonrandom mating within subpopulations (Fis) and <br />the effects of population subdivision (FST)" (Hartl and Clark 1997). <br />Table 2 shows the estimates of FIS over all loci for each sample. For purposes of <br />comparison, the values of Fis shown in parentheses for the upper Virgin River and Dexter <br />broodstock samples are the estimates calculated before the dataset was adjusted for null <br />allele frequencies. None of the samples had significant values of Fis (P > 0.05 after <br />correction for multiple tests) prior or subsequent to the adjustment for null allele <br />frequencies. These values indicate that within each population or sample, inbreeding is <br />minimal or nonexistent. <br />Genetic variation <br />Table 2 shows the results obtained using two measures of genetic variation: allelic <br />richness, and heterozygosity. The mean value of allelic richness (A) is an estimate of the <br />number of alleles per locus. Observed heterozygosity (Ho) is the frequency of observed <br />heterozygotes per sample; expected heterozygosity (HE) is the frequency of expected <br />11