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pattern than mottled sculpin, being most common in Browns Park, and were abundant there in <br />spring of each year (Fig. 38). Mountain whitefish was relatively abundant in spring 2003 seine <br />samples in Lodore Canyon, but rare in all other years, reaches, and seasons. Mountain whitefish <br />was most abundant in 2003, and we found a limited number as far downstream as Island- <br />Rainbow Park in spring, but few otherwise. Too few Colorado pikeminnow were captured in <br />seine samples to graph results; we discuss young pikeminnow captures in the species accounts <br />section. Only one bonytail, and no humpback chub or razorback sucker was captured in seine <br />samples. <br />Abundance patterns of the most common non-native fishes, red shiner and sand shiner, <br />generally increased in a downstream direction, being rare or absent in Browns Park, and <br />abundant downstream (Fig. 26, 27, 39). Fathead minnow was most common in upstream <br />Browns Park and declined in abundance downstream (Fig. 29, 40). Seasonal abundance patterns <br />of red shiner, sand shiner, and fathead minnow were highly variable, but generally increased <br />from spring to summer to autumn. Annual abundance was highest in 2002 and 2003 but lower in <br />2004. Higher spring abundance was not always associated with higher autumn abundance the <br />previous year. Green sunfish abundance was highest in Lodore Canyon and generally low <br />elsewhere (Fig. 40). In Lodore Canyon, spring and summer abundance was relatively low and <br />autumn abundance higher. Patterns elsewhere were variable. Redside shiner and white sucker <br />showed some consistencies in abundance patterns, being more common in upstream Browns <br />Park than in downstream reaches (Fig. 28, 30,41). White sucker showed seasonal abundance <br />patterns similar to native suckers, relatively low in spring, relatively high in summer, and lower <br />in autumn, and was most abundant in 2002 over all seasons, especially in Browns Park. Redside <br />shiner abundance was relatively high in spring and summer, but lower in autumn except in <br />Browns Park and Lodore Canyon in 2003, where autumn abundance was quite high. <br />Smallmouth bass were not found in Browns Park, but were more common in downstream <br />reaches (Fig. 42). Spring abundance of smallmouth bass was low, relatively high in summer, <br />and lower in autumn. Abundance was lowest in 2002 and higher in 2003 and 2004. <br />42