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In contrast to native fishes, non-native fishes generally remained abundant in the 2002 to <br />2004 period or increased in distribution and abundance in the study area since 1994 to 1996. Of <br />the five non-native species for which we compared electroshing CPUE data in Lodore Canyon <br />between the 1994 to 1996 and 2002 to 2004 periods, brown trout increased slightly and northern <br />pike remained similar in abundance between the two periods. Similarity in brown trout <br />abundance was somewhat surprising given the relatively warm conditions present during <br />summer in 2002 to 2004. During summer 2002, when water temperatures were warm, brown <br />trout were in poor condition and uncommon relative to 2003 and 2004. Brown trout apparently <br />quickly recolonized the reach from upstream and returned to former abundance levels. White <br />sucker, channel catfish, and smallmouth bass all increased in abundance in 2002 to 2004 <br />compared to the 1994 to 1996 period; the latter two taxa expanded their upstream distribution as <br />well. <br />Size distributions of those same five non-native fish captured by electrofishing all <br />changed in 2002 to 2004. Reasons for the bi-modal size distribution of brown trout in 2002 to <br />2004 were unknown, but may reflect reduced recruitment and abundance in 2002, a very warm <br />year. More large smallmouth bass and channel catfish were found upstream, perhaps suggesting <br />invasion from downstream rather than escapement from upstream Flaming Gorge Reservoir. <br />Age-0 smallmouth bass were detected only in lowermost Lodore Canyon in 2002 in low <br />numbers, the first documented reproduction there. More were noted in 2003 and 2004 and each <br />year moved progressively upstream. Reproduction was verified by collection of very small <br />smallmouth bass in drift net samples in 2003 and 2004, but not 2002, perhaps reflecting very low <br />reproduction in summer 2002. Smallmouth bass reproduction continues to expand in Lodore <br />Canyon, as an age-0 specimen was captured in autumn 2005 in uppermost Lodore Canyon. <br />Small white suckers were also more abundant upstream, possibly indicating higher recruitment <br />or higher reproduction there. <br />In Lodore Canyon, red shiner and sand shiner invaded upstream reaches in 2002 to 2004 <br />and were relatively more abundant there than in 1994 to 1996. Over all Lodore Canyon reaches, <br />relative abundance of those two taxa increased 925 and 352% respectively, and fathead minnow <br />36