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declined 81, 83, and 85%, respectively, between the periods. Mountain whitefish was the only <br />native fish to increase in abundance between the periods and that difference was slight. <br />Size structure changes for native fish in Lodore Canyon showed that small individuals <br />(300 mm TL or less) were less common in 2002 to 2004 compared to 1994 to 1996, which may <br />reflect diminished recruitment for those species. Reduced recruitment may also partially explain <br />lower abundance of adults in 2002 to 2004 period. Differences in roundtail chub length <br />frequency in the Green River upstream of the Yampa River (all large) and downstream (mixed <br />sizes) may reflect differences in the predator fish community in each reach; this is discussed <br />below in more detail. <br />Reduced absolute abundance of native fishes and lower recruitment in Lodore Canyon <br />and elsewhere may be a function of increased abundance of non-native fishes. For example, <br />increased abundance of red shiner may cause increased levels of predation on early life stages of <br />all native fishes, including catostomids (Ruppert et al. 1993, Bestgen et al. 1997, Bestgen et al. in <br />press). Fathead minnow is also an efficient predator on catostomid fish larvae, and their <br />increased abundance may be associated with reduced survival of early life stages of native fishes <br />(Dunsmoor 1993). Effects of small-bodied predators are likely most severe when age-1 and 2 <br />fish prey on larvae of native fishes. Larger larvae, juveniles, and small-bodied adults of some <br />native fishes that may or may not occupy backwaters may be affected by other predators as well. <br />These include expanding populations of smallmouth bass and channel catfish in the study area, <br />particularly Lodore Canyon, and brown trout which remained abundant in Lodore Canyon in <br />2002 to 2004. <br />Non-native fishes.-All trouts were combined to illustrate the general abundance pattern <br />for salmonids (excluding whitefish) because those cold-water taxa responded similarly to <br />environmental conditions. Brown trout constituted about 85% of all trouts captured, with <br />rainbow trout and rainbow x cutthroat trout hybrids (plus a few cutthroat trout, Snake River <br />subspecies) constituting the other 15%. Brown trout was found in all study reaches except <br />Island-Rainbow Park. In both time periods, salmonid abundance patterns were similar: they <br />were most common in upper Lodore Canyon (LD2) and were abundant (Fig. 24); CPUE for the <br />31