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<br />IIOI.IH ' :\.\:I) ST.\1.:\_\I\ER-SYSTI':,\I.\TICS of (;11,,1
<br />
<br />"Ill' lie 1\' 1I1l:lll1HT 11'0111 "1'1'1'1' (;1'1'1'11 Ri\'lT.
<br />.IIld Ihe all;,dlnl MlllCllI, 11'1' (llllllecled 10 Ihe
<br />d(".~"".\-rYf'}," (lllllpleX hy ,-ix ('OlllleniCllls,
<br />Fill: o[ 11,1'''' ICllllleniUls were 1'1'0111 IIIe
<br />"II;dl all;lI hed slIbdllsler 10 Ihe "eI('WIII.> X
<br />I YIJ},a" ".belllslel'; Ihc JlhCt, \\'as r"JIII Ihe
<br />'"lall ;lIlachnl slIhdlls el' 10 dIe "1-)'1"11/"
<br />,"hdIlSIIT. -This sllIall 'lIhdllSI<T of ciglll
<br />1III'IIIII('rs 1'(;I;ollle a hrid 'C I.elwcell Illc I\\'o
<br />1.'l'l;c (lllllpkxes, YCI il was joillcd 10 Ihc
<br />"lOll/HI,," (llISler hy (111)' 1\\'0 COllllcniollS.
<br />!lIlC fishrrolll lIppcr G 'CCII Ri\'cr illdlldctl
<br />ill thi, Inidgillg- slIhdll IeI' ill 1.c\'el 7 had
<br />IlIorc Ihall I" cOlllleniolS 10 Ihc "r"Il/I.\I,,"
<br />,Ir'sler alld was illdlld tl Ihel'c; Ihlls (111)'
<br />><:11'11 IIIclllhl'rs al'pcar d ill Ihc IJridgillg
<br />'""dllsler (Fi~, 3). Th l\I'lI lillkillg spcci,
<br />1IIl'IIS or 1.l'\c1 7 IJcI wc II Ihc "c)'I)III/" ami
<br />"c/cgaIH X cYIJlw" slIllL lISlcrs 1I0W Irad SC\'-
<br />n;ol 1II0rc ('OlllIcnioIlS 0 Ihe lauCl' gl'Olll'
<br />;lIId were illclllded Ihc c. :\Iso, OIlC [ish
<br />Ir"m I,ec's Fen)' alld 011 . [1'0111 f .ake I'o\\'ell
<br />111;11 cllleretl Ihe "dc!!.all,l" dllslcr ;11 l.c\'e1 5,
<br />;lIld IWo frCllll L ,ak., Pow'lI Ihal clllercd lire
<br />",-[e!:;,,"_," IllISl'T OIL 1.e 'c1 Ii. 1I0\\' I.elalllc
<br />IIhlrl' slrollgl)' (ollllened 10 Ihc "degrllH X
<br />'rfil/(/," slII.dllsler alld \\crc illdllded Iherc.
<br />\CI\' IlIcml.ITs cluslered illdllded. olle frolll
<br />1'1'1"'1' (;nTII \tilTr \\'illl ulle ((HI'":!:lioll 10
<br />1,,,,11 Lhc "de!!,,,",' X c)'IIIII" grollI' alld lhe
<br />,n'('II-III"IIII,cr InidgiHl:; S r1JCllISll'r; Olle fro\ll
<br />"/'I'IT Cn'('lI I{il'l'r all I 0111' frolll 1.ake
<br />1',,\\,1'11 ca, II wiLh a (0111 enioll 10 1.0111 lhc
<br />--q'/J/I"" c1usler alld lhe "e"'gans X /)'IJIW"
<br />,,,I,,IIISltT; ClIIC fish froll l.ake I '0 l\'l: 1 I 'I'ith
<br />1\\0 tOlllleniolls 10 lhe "deglllls" duster. 111'0
<br />"'"lIeniolls (0 lhe "clegllu X [ypllll" suh-
<br />II"SIlT. '"le 101lllenioll II Ihe "/')'/1/11/" suI.,
<br />tlllsler ;lJId OIlC IOlllleui HI 10 Ihe IJridgillg
<br />,,,I,, III'HT, Thesc fasL 0111' lish were ill-
<br />,I"dul ill l hc "dr'gllll,' 'TI)I/({" slIhdllslcr
<br />1,,1' ill I,CITI !I lhl')' Ilad IllOrc cOllllcnioll,'
<br />1'-' tltis gruup.
<br />l.e 1'<: 1 ~I,-Silllilaril)' -allle lowned 10
<br />illi,,-I:! I. This was tllC filial 1e1'e1 of Ihe
<br />I""gralll. Ihcrcfol-e. all s('clilllCIIS wen: iu,
<br />tI"d<:d_ Olle IICW IIICllrllCI fnllH ViI-gill RilTr
<br />1";111:<1 I hc .. ro Il//.I Ill" ell Slcr. Therc WCJ'<:
<br />,,"I)' six ('(Hllleniolls I.cI' 'Cell tllC "fobllsl,,"
<br />dllsler alld olher clllsters. The hrillgillg-
<br />,,,I,,III,lt'(' had 1'0111' n II IlcniOlls 10 tllC
<br />'/(/I"HI,," dllsler. ,\ lieII' IIlelllllcr dllslered
<br />"ilh lin: IOllllcniolls 10 lhe "cl".l!,IIII-'" dllslcr,
<br />1\\11 O'"II<TliOIlS 10 Ihe "1/ 1l//slll" dllslcr alld
<br />llllt CUIlIHO{ ';011 10 .he "('/t'P/IIl.\" X l.vIJI,ll"
<br />."1,, "I\lel. ()lIe lish 111111 "1'1"T (;IlTII
<br />
<br />115
<br />
<br />I{ inT 101l11<'llnl 10 Ille "C"'.l!,""_" X IYIJhll"
<br />slIl"IIISIIT alld Ihc "dt.'!!;IIII/' dllSICl' hy UIIC
<br />O'IIIICl'lioll 10 each. while a sc('ollll lish from
<br />11('('1'1' Crn'lI Ri,'cl' (,(lllllcoed UIII)' 10 Ihc
<br />ahu''C lie\\' IIIcIII1ICI'. These l\Yu lish \\'ere
<br />illdlllkd ill "c1cgllJ/.\ X cYIJlm" sulx:luslcr ill
<br />Fil-;' 3,
<br />
<br />C. f. foll//.I/1l awl (;. cl,'g'/JU'
<br />
<br />Thc laXilllel!"ics prugr;1I11 rc\'calcd littlc
<br />1I\(lrJ"mr..~' ,similaril)' lJclwcclI G. r.
<br />mfnlsf<< allli C. cleglllls. By Lc\'c1 2 of Ihe
<br />p....gram. H!J% o[ deglll/s alld 81,>,0 of ,-oul/.\la
<br />had 1IecII dllSlcrcll. This alld Ihc high simi-
<br />lalil)' ,-;dllc uf 1l.!11737 al Ihis Incl illdicalcd
<br />a high degrec of homog-elleily ill each [mm.
<br />Bolh dllslers of l.e\'e1 2 cOlllailled lIIelllhcrs
<br />1'1'0111 ;dl gCllcral geol:;ral'hical arcas ill ",hidl
<br />mUII.\I" ur c1eglll/s wcrc ('o\lcucd. 1;01' degllll_'
<br />Ihe ;Ireas wcre: uppcr GrcclI Ri,'cl-. Like
<br />Po\\'e1I, alld r .alc l\loh;l\'c. FOl' .-ollllSll1 Ihe
<br />alTas were: lippeI' GreclI Ri\u'. \\'hile River.
<br />IIPVcr Colorado Rivel'. Sail .Il1all Ri\'CI'. ;11,,1
<br />Black River of .-\rizoll;1. This sllggesls cadI
<br />1'01'\11 is III0rpholo~ic;rll)' IIl1iforlll 11......11:;'110111
<br />Ihe Colorado hasill, illllicalillg IWO dislillCl
<br />eWllllliollar}' lilies ralher lhall par;t1Ie1 cvolll-
<br />11011,
<br />llilTcrCIIlCS helwecn wU'Hla alld deglllls
<br />;'I'C sllllllllariLcd for lhe 19 characters IIscd ill
<br />Ihe laxillll'lrics ('rogr;1I11 ill Tahlc I. Thir-
<br />leell of I~I "holl" a dislilln sc('aLllioll helweell
<br />lhc I wo forlllS widl 1'0111' 011\(:1' c\r;,r;,ners
<br />showillg a lesser degrec o[ differellle, Ch;l1'-
<br />;'Clcrs I,e)[ sel'araling fOUIIJla ;lIld deg/II/J
<br />(1llalurc fish ~O(J nnn S1. or largel-) ;l1'C:
<br />dorsal alld all;rI fill ray tOlllllS. l-almslll !J-9.
<br />r/I'.~I/II,\' Ill-IO: head dCl'th. rolJ//.\11l 8li-I:!3,
<br />dr,~""S :.'J-Kti; "1'l'er j;"1' Iellglh, rubll"!/l
<br />K:l--117, dl'):.IlIl.\ :,ti--K-[; ;llld le;lSl dC('lh "I
<br />LllId;rI 1)(:(1,,,,, Ie, ",lno!1l 51--,,1, el,'gllll_"
<br />:!5---I!I, -'-he 1'0111' lI\cri-s1 ic char;lClers of - Cal,ie
<br />2 show hOlllogclleiL)' uf /'(Juusla and degllll.\
<br />IhrollghouL the Colorado uasill,
<br />V;lIIifel alld Kr;lIncr (191i!l) 1'0111,,1 sCl'eral
<br />life I,islory dillcrclllcs h"LlI"l'ell roll/Dill alld
<br />c!t....v,-ans whit II ~lllglllCIIl Illorpholllclric ~L"p~lr;l'
<br />lioll, Their sllId)' IT\'c;d('/1 difft-rcllces ill
<br />food hahils. r"IJlulll heillg lIIon: pist.i\'('HUJS
<br />l~au elCgllIlS; growlh rale. rolJ//JI/l (JIILtillg
<br />011 ils largesL i1llTC1I1C1I1 thc linl year o[ lil'c,
<br />d"K"",' ils fOllrlh year; alld Icllglh-weiglll
<br />relaliollshil's. r"ll/alll hCllJlllillg' he;,,'ier lhall
<br />dr',~'''I/S ;1S lellgLh ill<Teascs, Ecological dif-
<br />f<.'I"(" 1 1("('..... JWI\\T(,II 111l.....(' 1\\'(1 i()rlIIS ;In~ J"C-
<br />llet (, d I',clnn"d 1,.:llil;11. /(II,"-'O/U h
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