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<br />.1111
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<br />COPEI", Imll. NO. :1
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<br />(;. nlb,ls(n se/llil/lufn
<br />1'1 illcr (1!1.1li) examillcd the fi\'c Il'pc spet:i-
<br />mens of .l'eminnrla HllIcCled bl' Copc ami
<br />Yarrow (187:') alld temath'e1l' reg-;mled
<br />,w'/IIirllttfa to he a subspecies of G. I"OIIIuln.
<br />lie implied. as did Ellis (I!II1). thai lllis
<br />rorm was r""I<1 IhwIIghout the' Color;ulo
<br />hasill; hilt Ihe type material came fmm Vir-
<br />I)ill RivCJ". lIlah. Ellis HlIlsidered sl'Il/illl/tfa
<br />as specimells of /"Obl/sla with reduced squama-
<br />lioll 011 \'emral and dorsal areas. Some o[
<br />0111' specimens matdled this description, hut
<br />sillce they agree in all other char;lCters with
<br />Iypical mbll.l'(a, :Ire considered phenotypic
<br />\'arianls' for this one characH'.-. Therc(ore,
<br />we believe JClIlirl/{tfn is not a distinct elltitl'
<br />follnd Ihrollghout the Colorado hasill and
<br />th.at the name. sell/il/.lI~n rders to the Virgin
<br />RI\'er pOplll:ltlon o[ Czla.
<br />The Virgin River historically is an area
<br />o[ Jligh elldemism, suggesting a potential
<br />o[ dislillctiveness in the Gifn. populalion.
<br />Ollly six fish from Virgin Rh'Cr were t'X-
<br />amined, whidl gTeatly limil' conclusions.
<br />The laximelrics prog.'am placed the six [ish
<br />dose 10 robl/.f(n, J'el sOllletd':I! distillcl.
<br />Venehrae and gill r;lker ("(HIllIS (Tahle 2)
<br />are dislinn frOlll mbus(1/ or efrKl/lIs. l\liller's
<br />(/'U(i) data on the type sl'ecimellS ;lIso
<br />slIggested sell/illl/rln IIlay II(' dislillcl 11'0111
<br />r"hl/.I"(n. :\ddili,,"allll;Il<:rial from lhe Virgin
<br />RiH,[ is ncctkd hdore Ihe systemalic slallls
<br />o[ this poplllation call he UlldtTSlood.
<br />
<br />EI'OI.l!T10N
<br />Eml"tioll o[ Colorado hasill Gifa is
<br />IIl1dollllledly illteg-rally ticd \\'ith geologic;1I
<br />hislor'y o[ Ihat h;-tsi II , lIlIlortllllatclv, tltis
<br />hisl"r}' is preselllly ,sollletd':lc ohsCIlJT: The
<br />r"",,"'illl;' presc"tati,," aHCmpts to pr,,\'idc
<br />., ,'~""l'ral ",,,'illc of thc ('I'oJIIlioll of fishes
<br />\\'ii/lill ,he Colorado hasill. \I'i,1t spccial
<br />clIlphasis 011 Giia, IIsillg- the I;'eolog-ic illter.
<br />I"Tlatioll presellted ill J\/c/{ee d (fl. (19(;7).
<br />))urilll;' the Miocelle, the Cnlorado I'lateau
<br />was IIplifted ;11 III hC(';IIl1C higher thall Sill'.
<br />rOlllldillg Mea.s (Slllith. I'Hili; i\lcKec r.l /If,.
<br />I !Jri7), Tltis prohahly limited access to alld
<br />isol;lled the fishe.~ ill the hasill, i\lo,sl presclll
<br />1I;lIi,'e Colorado hasill fishes. espcci;"')' mill,
<br />1I0ll'S alld Slickers, probahly Irace their all.
<br />("estr}' to these fonlls. This helps explaill
<br />the hig-It deg-rec o[ elldemisrll foulld ill the
<br />hasill tOllay, Uplift of the K"ihah ll/,\\,;Ir/,
<br />di,i<l('(1 tlte hasill illtn 111"0 P;'I'lS, Th('
<br />11";''''p;li ))uill;lge S~Sl('1I1 he(,;III1(, ('st;d.,
<br />
<br />lished \\'est alld possihly SO III h west of Iht
<br />"1)\\'arp. alld Ihe .\1I1CSII',,1 Colorado Rin"
<br />dOlllill;<led Ihe uppcr hasill ill early,mid
<br />I'liocelle (McKee e( ai., 1!//i7). The Hllalap;li
<br />System was ch;lrancri~ed hy sltort. illll'ard
<br />draillage pallnlls ,,'hile 11t(' .\1I('('Slral (:01".
<br />rado Ri\'er \\'as Ihl'OlIgh,lIowillg ami ilia)
<br />kilT followed w/t;<l is 1I0W llle I.illle Colo.
<br />rado Ri,'er hasin ill a SOlllh(';Ist direni"n.
<br />The Ancestral Colorado Ri"er hecalllc
<br />pOllded in mid-Pliocelle ill nurtlteasterll
<br />;\riwna leavillg- the Bidahnclti FOllllalion
<br />Therefore. lhe Color;ldo ha.\ill ill e;lI'll'.ltIicl
<br />I'liocelle (Collsisted of '"'0 sl'p;,r;Ited s)';telll<
<br />The presellt Colorado ha.sill {ish fa IIn~
<br />reflens orig-in frolll all i,1t,.(.r",";\.ta a 10\\'('1'
<br />pan', the Grrrtnt Can)'on a~~.'neing tlte
<br />scpatatlllg pt)int; T!ljs.+Q{~~'I~,,~~
<br />edde...€C.~laIlY luwer h;lSllI elldemic fisltl"
<br />arc tYP;tally slll;dl riHT and strcalll [01'111<
<br />for exalllple se\Tral of the l'lagoplcrilli
<br />Cnlo,l'lo/llI/,f (i'(llIlo,l(nl.f) (1/1,.lri Baird ;11111
<br />(;;1';11'<1 alld (;iia l'oiJ//xla il/((,),I//{'''il/. SlIIilh
<br />(I!/(jli) sllggested tlt;1l C. da,.hi speci;lIed ill
<br />the lower hasill durillg Ihe I'liocelle ;111,1
<br />MilicI' alld Hu/,hs (/!/Iill) illdictted th;1t lit..
<br />"laW.ptl'I;II; also ("'011"(,11 ill Ihe 1'liot('I'"
<br />Tlte Ilualapai Systl'1II 11'011111 ),;"'e pro,id,,1
<br />all opportuuity lor SlIlh 101'11" 10 (,1..1"
<br />The hrge ril"('r ('III'ilOIIII"'1I1 01 "I{' "1'1'''
<br />hasill cOllld II;IIT {OllcIlITCIIII,. III'Ot';<I('(/ ,1
<br />placl' [or (:l"(tIUI;'"1 01 "large rilTr" ("1'111<
<br />Fossils o[ i'I)'dmd,,'illl,1 aud I ;il" lrolll ii"
<br />Bid;""Hhi (-'onll;t1iou (1.1)'l'II" alld ,\Iill<l.
<br />1!lli:i) that appear closely related LO 1'. 1/I{/lI'
<br />aud C. 1'. ruln/,Ifn alld/oJ' (;. d{'ga]/s alld,"1
<br />(;. (YI'"'' suggest I hcse larg(' l'ilTr [ish IILII
<br />halT had hegiullillgs ill the lippeI' I':,si"
<br />;\/illel' (1'/(;:') sUg~estl'd the Colorado 1I:"i"
<br />('l)'tI"J(.ftcilll.\' eHlln.II ill rhe l'liocelle. ,"llIi."
<br />(I ~H;fi) stlggt'sL<:d I he I;II-gc l'i\"Cl" Ca(o.\"/rnlllfl
<br />(f'al/(m/l'lIs) tliscol'(Jfllf Cope cH.hTd ill II"
<br />upper hasill dllrillg- the Pliocelle, 111('1('
<br />fore, :t tt<'O basiu hy\,olhesis of SP('('I:III'''I
<br />k" SOllie SIII'pOrt,
<br />;\1:111)' 1;lrge rileI' ronlls ;11(' I IITs, II I "
<br />II;"rihuled throughout Ihe Color;"lo I,,,,ill
<br />,\11 Cxpbllati(lII Iua)' be, II"h<"II the Col",;"I"
<br />KilTr .st;lIled 011 ils pJ'C,SOIl course t111""~/:
<br />Ihe Cralld Callyon it afforded a l'assag('I<;1I
<br />10 Ihe IOll'er hasill. ,This lat~ ri\'t'r ('/I
<br />\'ircmlllent \\'iJlIld Ita\'(' ;1( 1<'(1 as a /);urin I"
<br />the small strellll If'nlls 01' lhe l(ttl"{T '''1';''
<br />;llIeI 111;1,. c-xpbill e1isjllllll. di",illlll;'"'' "I
<br />\lId, ,"OI'IIIS :1\ 1'lfI.!!.fJ!J/r.'II\ {1/'.~l'l1f'_Hi",,,,\ (:1 TO
<br />"1111 r:, ria .I:i,
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