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-12- <br />high density shad population at Hite, in relation to.other lake areas, possibly <br />inhibited a secondary spawn. Secondary ccanplements of eggs either failed to <br />develop and/or latex developing younger females failed to produce a substantial <br />secondary spawning peak. <br />Numbers of shad captured correlated well with fish traces observed <br />in the epilimnion on sonar echograms (Figures 5, 6, and 7). Traces became <br />consistently mare numerous as the sampling season progressed. At Wahweap no <br />traces were observed on the June 20 echograms, and no fish were captured in <br />the trawl (Figure 5). On July 18 scattered traces were,observed, and 187 <br />shad were captured per tow. During September echograns showed dense groupings <br />of traces at all locations; however, trawl results were not consistently higher. <br />It has been shown that as shad grow their ability to avoid capture in a trawl <br />increases (Alfred Houser, personal catru iicaticn). Sonar echograms in August <br />and September represented mostly juvenile and adult shad.. The adults were <br />rarely captured, and juvenile catch decreased as size increased. Trawl catches <br />were probably biased toward smaller individuals in late summer. However, <br />decreased trawl catch in numbers and increased density of sonar fish traces <br />may not be totally inconsistent. Fewer but larger fish may cause denser fish <br />traces on sonar in cmparison to more numerous but smaller fish. Biomass was <br />generally highest in September, while shad numbers were below previous peaks <br />(Figures 3 and 4). Trawling depth may also have been a factor contributing <br />to reduced catch in Septenber. Maximimi depth of the standard tow may not have <br />been deep enough to sample fish in the 10-12 m range (35-40 ft) as shown by <br />Comparative trawl samples for 1976 and 1977 showed a stronger year <br />class of shad in Wahweap Bay (Table 1). Others have noticed relatively <br />large differences in year class strength of shad (Bryant and Houser 1968,