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(~ Average length of the frost-free season (V6~. The length of the frost- <br />free season correlates with the length of the growing season for northern <br />pike; however, if the cold weather season is too short, normal gonadal devel- <br />opment may be impaired. The suitability index curve is based on growth data <br />for northern pike, laboratory observations on yellow perch spawning (see <br />text), the relative position of isopleths for length of frost free season <br />(Visher 1954), and the southern limit to the native range of northern pike <br />(Grossman 1978). <br />Maximal weekly average temperature of surface water (1 to 2 m deep) (V,~. <br />Summer water temperature affects growth and survival of northern pike. The <br />optimal temperature for growth of fingerling and older northern pike is 19 to <br />20° C (Casselman .1978). Temperatures will be in the optimal range during <br />warming and cooling phases if the maximal summer temperature is slightly <br />higher than the growth optimum. Maximal suitability of V, is assumed for <br />peak weekly average temperatures of 20 to 25° C. Laboratory studies show that <br />temperatures over 32° C can kill northern pike within several days (Scott <br />1964). An oxygenated metalimnion, mouths of tributaries or inlets, or springs <br />may be used when temperatures elsewhere are too high. Their effectiveness as <br />thermal refugia depend on their size and the duration of hot spells. <br />Percent pools and backwaters during midsummer (U$) and stream gradient <br />,(.V9~. Riverine habitat may be suitable to the extent that it resembles a lake <br />,~ environment. Northern pike require standing, or very slowly moving, water. <br />Two variables are included as indicators of the presence of low velocity <br />habitat: percent pools and backwaters during midsummer and stream gradient. <br />A direct, proportional relationship between the availability of standing water <br />habitat (V$} and habitat suitability is assumed. However, a high gradient <br />stream is likely to be of low suitability even if pools are present, because <br />movement between pools would be restricted. It is assumed that northern pike <br />are less well adapted to high gradient streams than are muskellunge, based on <br />patterns of distribution for the two species (Grossman 1978; Harrison and <br />Hadley 1978). Muskellunge occur in stream reaches with gradients as high as <br />6.9 mlkm (Parsons 1959), but gradients less than 2 m/km seem to be more typical <br />of riverine muskellunge habitat (Brewer 1970; Miles 1978). It is assumed that <br />stream reaches with gradients greater than 5 m/km (0.5%) have little or no <br />habitat value for northern pike. <br />Field Use of the Model <br />Year to year variation in weather complicates the application of this <br />habitat model. Such variability is most pronounced for variables related to <br />temperature (V6 and V,) and water level (U1 and V2). Between-year variation <br />does not pose a problem if the objective of an HSI model application is to <br />obtain an HSI rating for a single year. More often, however, the goal is to <br />obtain an average or representative HSI for a particular body of water that <br />can be compared with HSI`s for other bodies of water or' for the same body of <br />25 <br />