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72,000 fingerlings/ha), both among marshes and among years. The median <br />(2,717 fingerlings/ha) was used in the following calculations. <br />Survival of. fingerlings through adulthood is also highly variable. <br />Estimated survival of fingerling northern pike through their first summer was <br />6%, 22%, and 63% in three successive years (mean = 28°0) in a Michigan lake <br />(Beyerle and Williams 1972). Assuming 65°o annual mortality thereafter [which <br />seems reasonable for lakes in the conterminous United States, on the basis of <br />data compiled by Carlander (1970)], 9.8% of the fingerlings would survive to <br />age 1+, 3.4°o to age 2+, 1.2% to age 3+, and 0.4% to age 4+. Ages 2 through 4 <br />usually account for most of the breeding stock for populations in the conter- <br />minous U.S. Assuming a stable age structure and that most mortality occurs <br />between autumn and spring, there would be a total of 0.034 + 0.012 + 0.004 = <br />0.050 adult fish in the population per fingerling produced in spawning habitat. <br />The minimum ratio of spawning to summer habitat can be calculated in the <br />following way: <br />spawning habitat area _ max. density of adults x 1 <br />summer habitat area spawning habitat productivity <br />x fingerlings per adult <br />= 40 adults/ha summer habitat <br />x 1 ha spawning habitat/2,717 fingerlings <br />~~ x 1 fingerling/0.05 adults <br />= 0.29 ha spawning habitat/ha summer habitat <br /> <br />If it is assumed that most mortality .occurs between spring and autumn, <br />then the 1+ fish should be included in the calculation. There would be <br />0.098 + 0.034 + 0.012 + 0.004 = 0.148 adult fish in the population/fingerling <br />produced, and the minimum ratio of spawning to summer habitat would be 0.10. <br />Both values are similar to that obtained using the method based on spawn- <br />ing behavior (0.40). An intermediate value, 0.25, is used to develop a suit- <br />ability index curve. It is assumed that optimal northern pike habitat must <br />have at least 0.25 ha of spawning habitat for each ha of summer habitat. <br />Habitats in which the relative availability of spawning habitat is considerably <br />less than this can still have high suitability index ratings for V1 because of <br />the asymptotic shape of the suitability index curves. <br />Type of vegetation in spawning areas is also important. The probability <br />that fertilized eggs will settle on vegetation as they sink depends on vegeta- <br />tion density. Young northern pike also depend on vegetation for protective <br />cover and on the associated invertebrate fauna for food. Lush vegetative <br />cover is ideal; sparse cover is poor. The suitability index for V1 can be <br />maximal (1.0) only if proper vegetation is present. <br />23 <br />