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The model is designed to assign the highest HSI's to systems capable of <br />producing the most northern pike biomass per unit time on a sustained basis, <br />regardless of how that biomass is apportioned among individual fish. Few long <br />term data are available on production, yield, standing crop, or other measures <br />of population performance for northern pike under different environmental <br />conditions. Therefore, behavioral observations and measures of individual <br />performance, such as growth and survival, were also used to develop suitability <br />index curves for model variables. The rationale and assumptions used to <br />develop the curves are presented in a following section. <br />Modifications to the model, such as including different or additional <br />variables, may be necessary in some situations. The model is structurally <br />.simple, and madifications are easily made. <br />Model Applicability <br />Geographic area. The model is applicable to lakes, reservoirs, rivers, <br />and streams throughout North America. <br />Season. The model is structured to account for seasonal changes in <br />weather and habitat requirements of northern pike and, thus, to indicate the <br />ability of a'-given habitat to sustain a population on a year-round basis. <br />Overwintering requirements may not be adequately addressed for northern pike <br />in shallow lakes~or low discharge rivers that experience extended ice cover <br />(see below). <br />Cover types. The model is applicable to permanent lakes, ponds, and <br />reservoirs with a mean depth _> 1 m during midsummer. It is of doubtful reli- <br />ability .for shallow lakes (even those with mean depths > 1 m) that are covered <br />with ice for long periods. The suitability index graph for V3 (curve A) is <br />the only part of the model-that addresses the possibility of winterkill of <br />northern pike due to low dissolved oxygen concentrations. It is recognized <br />that any single variable is an inadequate indicator of the occurrence and <br />severity of this phenomenon. However, an accurate, general model for predict- <br />ing winter dissolved oxygen concentrations, based on easily measured or <br />predicted habitat characteristics, does not exist. [The empirical techniques <br />developed by Barica and Mathias (1979) for predicting winterkill in shallow <br />prairie lakes may be useful in some cases.] Because some of the factors that <br />affect the likelihood of winterkill are highly variable from year to year, <br />dissolved oxygen measurements for a single winter do not -adequately indicate <br />the likelihood of occurrence of winterkill in a given lake over a series of <br />years. The fact that northern pike can find and use pockets or layers of <br />water with relatively high oxygen concentrations (Magnuson and Karlen 1970; <br />Scidmore 1970} further complicates the modeling problem. Habitats that <br />experience occasional, partial winterkill of northern pike may nonetheless be <br />highly productive of northern pike. Rather than include additional variables <br />with no guarantee of improved model accuracy, it was decided to keep the model <br />simple and emphasize its questionable applicability to potential winterkill <br />lakes. Information presented by Greenbank (1945) and Schneberger (1970) <br />15 <br />